Comparison of 3 files, rendered from file:/u:/diff/diff-diff.xml

Comparison generated August 4, 2020, 6:31 A.M.
All sluglines were removed before versions were compared.
Differences
URILengthNumberLengthPortion
darwin_1859_01
Tt
Tt
file:/u:/darwin/darwin_1859_01.xml69066962832812.1%
darwin_1860_01
Tt
Tt
file:/u:/darwin/darwin_1860_01.xml69331964859312.4%
darwin_1872_01
Tt
Tt
file:/u:/darwin/darwin_1872_01.xml735149501277617.4%
file:/u:/diff/diff-diff.xml8199919192126125.9%
Similarity of pairs of files (%)
The table below shows the percentage of similarity of each pair of versions, starting with the version that shows the least divergence from the entire group and proceeding to versions that are most divergent. This table is useful for identifying clusters and pairs of versions that are closest to each other.
darwin_1859_01darwin_1860_01darwin_1872_01
darwin_1859_0199.385.4
darwin_1860_0199.385.5
darwin_1872_0185.485.5
Divergence of sets of three files (string-length)
Versions are presented below in sets of three, with a Venn diagram for visualization. Numbers refer to the quantity of characters that diverge from common, shared text (that is, shared by all three, regardless of any other version).
The diagrams are useful for thinking about how a document was revised. For example, if you know that versions A, B, and C represent consecutive editing stages in a document, then focus on the circle for B. Its relative distance from A and C indicate how far along the editing process it went. The diagrams also depict wasted work. Whatever is in B that is in neither A nor C represents text that B added that C deleted. Whatever is in A and C but not in B represent text deleted by B that was restored by C.
Note, some data combinations are impossible to draw accurately with a 3-circle Venn diagram (e.g., a 3-circle Venn diagram for items in the set {[a, z], [b, z], [c, z]} will always mistakenly show overlap for each pair of items).
The colors are fixed according to the A, B, and C components of the Venn diagram, not to the version labels, which will change color from one Venn diagram to the next.
darwin_1859_01
darwin_1860_01
darwin_1872_01
The distance from
darwin_1859_01
to
darwin_1872_01
is 20886 (8219 characters deleted and 12667 inserted). Intermediate version
darwin_1860_01
contributed 552 characters to the end result (2.6%). But it inserted 266 characters that were deleted by
darwin_1872_01
, and deleted 109 characters that were restored by
darwin_1872_01
. Nixed insertions:
(2),
i
(2),
ar
(1),
or
(1),
some
(1),
the␠breeds␠h
(1),
v
(1),
e
(1),
.␠Knowing,␠as␠we␠do,␠that␠savages␠are␠very␠fond
(1),
taming␠
(1),
mals,␠it␠seems␠to␠me␠unlikely,␠in␠the␠case␠of␠the␠
(1),
og-genus,␠which␠is␠distributed␠in␠a␠wild␠state␠throughout␠the␠world,␠that␠since␠man␠first␠appeared␠one␠single␠species␠alone␠should
(1),
at
(1),
va
(1),
-
(1),
z
(1). Nixed deletions:
'
(5),
,
(1),
the␠s
(1),
me
(1),
-
(1),
which␠are␠first␠given␠to␠him␠in␠some␠slight␠degree␠by␠nature.␠No␠man␠would␠ever␠try␠to␠make␠a␠
(1),
s
(1).
Aggregate progress was 182 (0.9%, after adjusting for 5 nixed deletions and insertions that seem negligible).
*To show more accurately the differences between the three versions, the proportionate size of the central common section has been reduced by 48401, to match the size of the largest sliver. All other non-common slivers are rendered proportionate to one another.

Comparison

darwin_1859_01 darwin_1860_01 darwin_1872_01
WHEN we
darwin_1859_01 darwin_1860_01
l
darwin_1872_01
c
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
ok to
darwin_1872_01
mpare
darwin_1859_01 darwin_1860_01 darwin_1872_01
the individuals of the same variety or sub-variety of our older cultivated plants and animals, one of the first points which strikes us
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
is, that they generally differ m
darwin_1859_01
uch m
darwin_1859_01 darwin_1860_01 darwin_1872_01
ore from each other
darwin_1859_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
than do the individuals of any one species or variety in a state of nature.
darwin_1859_01 darwin_1860_01
Whe
darwin_1872_01
A
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1872_01
d if
darwin_1859_01 darwin_1860_01 darwin_1872_01
we reflect on the vast diversity of the plants and animals which have been cultivated, and which have varied during all ages under the most different climates and treatment,
darwin_1859_01 darwin_1860_01
I think
darwin_1859_01 darwin_1860_01 darwin_1872_01
we are driven to conclude that this great
darwin_1859_01
er
darwin_1859_01 darwin_1860_01 darwin_1872_01
variability is
darwin_1859_01 darwin_1860_01
simply
darwin_1859_01 darwin_1860_01 darwin_1872_01
due to our domestic productions having been raised under conditions of life not so uniform as, and somewhat different from, those to which the parent-species ha
darwin_1859_01 darwin_1860_01
ve
darwin_1872_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
been exposed under nature. There is
darwin_1859_01 darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
also,
darwin_1859_01 darwin_1860_01
I think,
darwin_1859_01 darwin_1860_01 darwin_1872_01
some probability in the view propounded by Andrew Knight, that this variability may be partly connected with excess of food. It seems
darwin_1859_01 darwin_1860_01
pretty
darwin_1859_01 darwin_1860_01 darwin_1872_01
clear that organic beings must be exposed during several generations to
darwin_1859_01 darwin_1860_01
the
darwin_1859_01 darwin_1860_01 darwin_1872_01
new conditions
darwin_1859_01 darwin_1860_01
of life
darwin_1859_01 darwin_1860_01 darwin_1872_01
to cause any
darwin_1859_01 darwin_1860_01
app
darwin_1872_01
g
darwin_1859_01 darwin_1860_01 darwin_1872_01
re
darwin_1859_01 darwin_1860_01
ci
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
ble
darwin_1872_01
t
darwin_1859_01 darwin_1860_01 darwin_1872_01
amount of variation; and that
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
when the organisation has once begun to vary, it generally continues
darwin_1859_01 darwin_1860_01
to
darwin_1859_01 darwin_1860_01 darwin_1872_01
vary
darwin_1872_01
ing
darwin_1859_01 darwin_1860_01 darwin_1872_01
for many generations. No case is on record of a variable
darwin_1859_01 darwin_1860_01
be
darwin_1872_01
organ
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
ng
darwin_1872_01
sm
darwin_1859_01 darwin_1860_01 darwin_1872_01
ceasing to
darwin_1859_01 darwin_1860_01
be
darwin_1859_01 darwin_1860_01 darwin_1872_01
var
darwin_1859_01 darwin_1860_01
iable
darwin_1872_01
y
darwin_1859_01 darwin_1860_01 darwin_1872_01
under cultivation. Our oldest cultivated plants, such as wheat, still
darwin_1859_01 darwin_1860_01
often
darwin_1859_01 darwin_1860_01 darwin_1872_01
yield new varieties: our oldest domesticated animals are still capable of rapid improvement or modification.
darwin_1859_01 darwin_1860_01
It
darwin_1872_01
As
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
h
darwin_1872_01
far
darwin_1859_01 darwin_1860_01 darwin_1872_01
as
darwin_1872_01
I am a
darwin_1859_01 darwin_1860_01 darwin_1872_01
b
darwin_1872_01
l
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
to judg
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
n
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
dispu
darwin_1872_01
af
darwin_1859_01 darwin_1860_01 darwin_1872_01
te
darwin_1859_01 darwin_1860_01
d
darwin_1872_01
r long
darwin_1859_01 darwin_1860_01 darwin_1872_01
at
darwin_1872_01
tending
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
w
darwin_1872_01
to t
darwin_1859_01 darwin_1860_01 darwin_1872_01
h
darwin_1859_01 darwin_1860_01
a
darwin_1872_01
e subjec
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
p
darwin_1872_01
th
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
r
darwin_1872_01
condit
darwin_1859_01 darwin_1860_01 darwin_1872_01
io
darwin_1859_01 darwin_1860_01
d
darwin_1872_01
ns
darwin_1859_01 darwin_1860_01 darwin_1872_01
of life
darwin_1872_01
appear to act in two ways,—directly on the whole organisation or on certain parts alone, and indirectly by affecting the reproductive system. With respect to the direct action, we must bear in mind that in every case, as Professor Weismann has lately insisted, and as I have incidentally shown in my work on 'Variation under Domestication,' there are two factors: namely, the nature of the organism, and the nature of the conditions. The former seems to be much the more important; for nearly similar variations sometimes arise under, as far as we can judge, dissimilar conditions; and, on the other hand, dissimilar variations arise under conditions which appear to be nearly uniform. The effects on the offspring are either definite or indefinite. They may be considered as definite when all or nearly all the offspring of individuals exposed to certain conditions during several generations are modified in the same manner. It is extremely difficult to come to any conclusion in regard to the extent of the changes which have been thus definitely induced. There can, however, be little doubt about many slight changes,—such as size from the amount of food, colour from the nature of the food, thickness of the skin and hair from climate, &c. Each of the endless variations which we see in the plumage of our fowls must have had some efficient cause; and if the same cause were to act uniformly during a long series of generations on many individuals, all probably would be modified in the same manner. Such facts as the complex and extraordinary out-growths which invariably follow from the insertion of a minute drop of poison by a gall-producing insect, show us what singular modifications might result in
darwin_1859_01 darwin_1860_01 darwin_1872_01
the ca
darwin_1859_01 darwin_1860_01
u
darwin_1859_01 darwin_1860_01 darwin_1872_01
se
darwin_1859_01 darwin_1860_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
of
darwin_1872_01
plants from a chemical change in the nature of the sap. Indefinite
darwin_1859_01 darwin_1860_01 darwin_1872_01
variability
darwin_1872_01
is a much more common result of changed conditions than definite variability
darwin_1859_01 darwin_1860_01 darwin_1872_01
,
darwin_1859_01 darwin_1860_01
w
darwin_1872_01
and
darwin_1859_01 darwin_1860_01 darwin_1872_01
ha
darwin_1872_01
s probably played a more impor
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
eve
darwin_1872_01
ant pa
darwin_1859_01 darwin_1860_01 darwin_1872_01
r
darwin_1872_01
t in
darwin_1859_01 darwin_1860_01 darwin_1872_01
the
darwin_1859_01 darwin_1860_01
y
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1872_01
for
darwin_1859_01 darwin_1860_01 darwin_1872_01
ma
darwin_1859_01 darwin_1860_01
y be, generally ac
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
;
darwin_1872_01
ion
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
whethe
darwin_1872_01
of ou
darwin_1859_01 darwin_1860_01 darwin_1872_01
r d
darwin_1859_01 darwin_1860_01
u
darwin_1872_01
omestic races. We see indefinite va
darwin_1859_01 darwin_1860_01 darwin_1872_01
ri
darwin_1872_01
ability i
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1859_01 darwin_1860_01
g
darwin_1859_01 darwin_1860_01 darwin_1872_01
the e
darwin_1859_01 darwin_1860_01
ar
darwin_1872_01
nd
darwin_1859_01 darwin_1860_01 darwin_1872_01
l
darwin_1859_01 darwin_1860_01
y
darwin_1872_01
ess
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
or
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
l
darwin_1859_01 darwin_1860_01
a
darwin_1872_01
igh
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
pe
darwin_1872_01
culia
darwin_1859_01 darwin_1860_01 darwin_1872_01
ri
darwin_1859_01 darwin_1860_01
od
darwin_1872_01
ties
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
of
darwin_1872_01
which
darwin_1859_01 darwin_1860_01 darwin_1872_01
d
darwin_1872_01
istinguish th
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
indi
darwin_1859_01 darwin_1860_01 darwin_1872_01
v
darwin_1859_01 darwin_1860_01
e
darwin_1872_01
idua
darwin_1859_01 darwin_1860_01 darwin_1872_01
l
darwin_1859_01 darwin_1860_01
opment
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
of the
darwin_1872_01
sam
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
species, and which cannot be accounted for by inheritance fro
darwin_1859_01 darwin_1860_01 darwin_1872_01
m
darwin_1859_01 darwin_1860_01
b
darwin_1872_01
eithe
darwin_1859_01 darwin_1860_01 darwin_1872_01
r
darwin_1859_01 darwin_1860_01
yo,
darwin_1872_01
parent
darwin_1859_01 darwin_1860_01 darwin_1872_01
or
darwin_1872_01
from some more remote
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1872_01
nces
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1872_01
or. Even strongly-marked differences occasionally appear in
darwin_1859_01 darwin_1860_01 darwin_1872_01
the
darwin_1859_01 darwin_1860_01
instant of conception. Geoffro
darwin_1859_01 darwin_1860_01 darwin_1872_01
y
darwin_1859_01 darwin_1860_01
St. Hilaire's experiments sh
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
w that
darwin_1859_01 darwin_1860_01 darwin_1872_01
un
darwin_1859_01 darwin_1860_01
natural treatment
darwin_1872_01
g
darwin_1859_01 darwin_1860_01 darwin_1872_01
of the
darwin_1872_01
sam
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
litter, and in seedlings fro
darwin_1859_01 darwin_1860_01 darwin_1872_01
m
darwin_1859_01 darwin_1860_01
b
darwin_1872_01
the same seed-capsule. At long intervals of time, out of millions of individuals reared in the same count
darwin_1859_01 darwin_1860_01 darwin_1872_01
ry
darwin_1872_01
and fed
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1872_01
n nearly the same food, deviations of structure so strongly pronounced as to deserve to be
darwin_1859_01 darwin_1860_01 darwin_1872_01
ca
darwin_1859_01 darwin_1860_01
us
darwin_1872_01
ll
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
s
darwin_1872_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
monstrosities
darwin_1872_01
arise
darwin_1859_01 darwin_1860_01 darwin_1872_01
;
darwin_1859_01 darwin_1860_01
and
darwin_1872_01
but
darwin_1859_01 darwin_1860_01 darwin_1872_01
monstrosities cannot be separated by any
darwin_1859_01 darwin_1860_01
clear line of
darwin_1859_01 darwin_1860_01 darwin_1872_01
distinct
darwin_1872_01
l
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
o
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1872_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
from
darwin_1859_01 darwin_1860_01
m
darwin_1872_01
slight
darwin_1859_01 darwin_1860_01 darwin_1872_01
er
darwin_1859_01 darwin_1860_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
variations.
darwin_1859_01 darwin_1860_01
B
darwin_1872_01
All s
darwin_1859_01 darwin_1860_01 darwin_1872_01
u
darwin_1872_01
ch changes of structure, whether extremely sligh
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
I am
darwin_1872_01
or
darwin_1859_01 darwin_1860_01 darwin_1872_01
strongly
darwin_1859_01 darwin_1860_01
inclin
darwin_1872_01
mark
darwin_1859_01 darwin_1860_01 darwin_1872_01
ed
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
to
darwin_1872_01
which
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
sus
darwin_1872_01
ap
darwin_1859_01 darwin_1860_01 darwin_1872_01
pe
darwin_1859_01 darwin_1860_01
ct th
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
t
darwin_1872_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
the
darwin_1872_01
a
darwin_1859_01 darwin_1860_01 darwin_1872_01
mo
darwin_1872_01
ng
darwin_1859_01 darwin_1860_01 darwin_1872_01
st
darwin_1859_01 darwin_1860_01
frequent c
darwin_1872_01
m
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
use
darwin_1872_01
ny
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
of var
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1872_01
ndividu
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
bi
darwin_1872_01
ls
darwin_1859_01 darwin_1860_01 darwin_1872_01
li
darwin_1872_01
ving
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
y
darwin_1872_01
ogether,
darwin_1859_01 darwin_1860_01 darwin_1872_01
may be
darwin_1859_01 darwin_1860_01
attributed to the male and female reproductive elements having been affected prior to the act of
darwin_1859_01 darwin_1860_01 darwin_1872_01
con
darwin_1859_01 darwin_1860_01
cept
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
on. Sev
darwin_1872_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
er
darwin_1859_01 darwin_1860_01
al r
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
asons
darwin_1872_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
m
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
ke me believe in this; but the chief one i
darwin_1859_01 darwin_1860_01 darwin_1872_01
s the
darwin_1859_01 darwin_1860_01
r
darwin_1872_01
ind
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
markabl
darwin_1872_01
finit
darwin_1859_01 darwin_1860_01 darwin_1872_01
e effect
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
w
darwin_1872_01
of t
darwin_1859_01 darwin_1860_01 darwin_1872_01
h
darwin_1859_01 darwin_1860_01
ich
darwin_1872_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
con
darwin_1872_01
ditions of life on each individual organism, in nearly the same manner as a chill affects different men in an inde
darwin_1859_01 darwin_1860_01 darwin_1872_01
fin
darwin_1872_01
it
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
m
darwin_1872_01
ann
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
r, accordi
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1872_01
g
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1872_01
o their state of body
darwin_1859_01 darwin_1860_01 darwin_1872_01
or c
darwin_1859_01 darwin_1860_01
ul
darwin_1872_01
ons
darwin_1859_01 darwin_1860_01 darwin_1872_01
ti
darwin_1859_01 darwin_1860_01
v
darwin_1872_01
tution, causing coughs or colds, rheumatism, or inflamm
darwin_1859_01 darwin_1860_01 darwin_1872_01
ation
darwin_1872_01
s of various organs. With respect to what I
darwin_1859_01 darwin_1860_01 darwin_1872_01
ha
darwin_1859_01 darwin_1860_01
s
darwin_1872_01
ve
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
on
darwin_1872_01
called
darwin_1859_01 darwin_1860_01 darwin_1872_01
the
darwin_1859_01 darwin_1860_01
fu
darwin_1872_01
i
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1872_01
direct a
darwin_1859_01 darwin_1860_01 darwin_1872_01
ction
darwin_1859_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
of
darwin_1872_01
changed conditions, namely, through
darwin_1859_01 darwin_1860_01 darwin_1872_01
the reproductive system
darwin_1859_01 darwin_1860_01
;
darwin_1872_01
being affected, we may infer that variability is thus induced, partly from the fact of
darwin_1859_01 darwin_1860_01 darwin_1872_01
this system
darwin_1859_01 darwin_1860_01
app
darwin_1872_01
b
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
ar
darwin_1859_01 darwin_1860_01 darwin_1872_01
ing
darwin_1859_01 darwin_1860_01
to b
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
far mo
darwin_1872_01
xt
darwin_1859_01 darwin_1860_01 darwin_1872_01
re
darwin_1872_01
mely
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1859_01 darwin_1860_01
usc
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
p
darwin_1872_01
nsi
darwin_1859_01 darwin_1860_01 darwin_1872_01
ti
darwin_1859_01 darwin_1860_01
bl
darwin_1872_01
v
darwin_1859_01 darwin_1860_01 darwin_1872_01
e t
darwin_1859_01 darwin_1860_01
han any
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
ther part of the organisation, to the action of
darwin_1859_01 darwin_1860_01 darwin_1872_01
any change in the conditions
darwin_1872_01
, and partly from the similarity, as Kölreuter and others have remarked, between the variability which follows from the crossing
darwin_1859_01 darwin_1860_01 darwin_1872_01
of
darwin_1872_01
distinct species, and that which may be observed with plants and animals when reared under new or unnatural conditions. Many facts clearly show how eminently susceptible the reproductive system is to very s
darwin_1859_01 darwin_1860_01 darwin_1872_01
li
darwin_1859_01 darwin_1860_01
f
darwin_1872_01
ght chang
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
s in the surrounding conditions
darwin_1859_01 darwin_1860_01 darwin_1872_01
. Nothing is more easy than to tame an animal, and few things more difficult than to get it to breed freely under confinement, even
darwin_1859_01 darwin_1860_01
in the many cases
darwin_1859_01 darwin_1860_01 darwin_1872_01
when the male and female unite. How many animals there are which will not breed, though
darwin_1859_01 darwin_1860_01
living long und
darwin_1872_01
k
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
r no
darwin_1872_01
p
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
very
darwin_1872_01
in
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
c
darwin_1872_01
an a
darwin_1859_01 darwin_1860_01 darwin_1872_01
l
darwin_1872_01
m
darwin_1859_01 darwin_1860_01 darwin_1872_01
os
darwin_1872_01
t fre
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
confinemen
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1872_01
ate
darwin_1859_01 darwin_1860_01 darwin_1872_01
in their native country! This is generally
darwin_1872_01
, but erroneously,
darwin_1859_01 darwin_1860_01 darwin_1872_01
attributed to vitiated instincts
darwin_1859_01 darwin_1860_01
;
darwin_1872_01
.
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
but how m
darwin_1872_01
M
darwin_1859_01 darwin_1860_01 darwin_1872_01
any cultivated plants display the utmost vigour, and yet rarely or never seed! In some few
darwin_1859_01 darwin_1860_01
such
darwin_1859_01 darwin_1860_01 darwin_1872_01
cases it has been
darwin_1860_01 darwin_1872_01
disc
darwin_1859_01
f
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1860_01 darwin_1872_01
vere
darwin_1859_01
un
darwin_1859_01 darwin_1860_01 darwin_1872_01
d
darwin_1859_01
out
darwin_1859_01 darwin_1860_01 darwin_1872_01
that
darwin_1872_01
a
darwin_1859_01 darwin_1860_01 darwin_1872_01
very trifling change
darwin_1859_01 darwin_1860_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
, such as a little more or less water at some particular period of growth, will determine whether or not
darwin_1859_01 darwin_1860_01
the
darwin_1872_01
a
darwin_1859_01 darwin_1860_01 darwin_1872_01
plant
darwin_1859_01 darwin_1860_01
s
darwin_1872_01
will produc
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
ts a
darwin_1859_01 darwin_1860_01 darwin_1872_01
seed
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
. I cannot here
darwin_1872_01
giv
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
nter on
darwin_1859_01 darwin_1860_01 darwin_1872_01
the
darwin_1859_01 darwin_1860_01
copious
darwin_1859_01 darwin_1860_01 darwin_1872_01
details which I have collected
darwin_1872_01
and elsewhere published
darwin_1859_01 darwin_1860_01 darwin_1872_01
on this curious subject; but to show how singular the laws are which determine the reproduction of animals under confinement, I may
darwin_1859_01 darwin_1860_01
just
darwin_1859_01 darwin_1860_01 darwin_1872_01
mention that carnivorous animals, even from the tropics, breed in this country pretty freely under confinement, with the exception of the plantigrades or bear family
darwin_1872_01
, which seldom produce young
darwin_1859_01 darwin_1860_01 darwin_1872_01
; whereas
darwin_1859_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
carnivorous birds, with the rarest exceptions, hardly ever lay fertile eggs. Many exotic plants have pollen utterly worthless, in the same
darwin_1859_01 darwin_1860_01
exact
darwin_1859_01 darwin_1860_01 darwin_1872_01
condition as in the most sterile hybrids. When, on the one hand, we see domesticated animals and plants, though often weak and sickly,
darwin_1859_01 darwin_1860_01
yet
darwin_1859_01 darwin_1860_01 darwin_1872_01
breeding
darwin_1859_01 darwin_1860_01
quite
darwin_1859_01 darwin_1860_01 darwin_1872_01
freely under confinement; and when, on the other hand, we see individuals, though taken young from a state of nature
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
perfectly tamed, long-lived
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
and healthy (of which I could give numerous instances), yet having their reproductive system so seriously affected by unperceived causes as to fail
darwin_1859_01 darwin_1860_01
in
darwin_1872_01
to
darwin_1859_01 darwin_1860_01 darwin_1872_01
act
darwin_1859_01 darwin_1860_01
ing
darwin_1859_01 darwin_1860_01 darwin_1872_01
, we need not be surprised at this system, when it does act under confinement, acting
darwin_1859_01 darwin_1860_01
not qu
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
te
darwin_1872_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
regularly, and producing offspring
darwin_1859_01 darwin_1860_01
n
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1872_01
mewha
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
perfectly
darwin_1872_01
un
darwin_1859_01 darwin_1860_01 darwin_1872_01
like their parents
darwin_1859_01
or variable
darwin_1859_01 darwin_1860_01 darwin_1872_01
.
darwin_1859_01 darwin_1860_01
Sterility has been said to be the bane of horticulture; but on this view we owe variability to the same cause which produces sterility; and variability is the source of all the choicest productions of the garden.
darwin_1859_01 darwin_1860_01 darwin_1872_01
I may add, that as some organisms
darwin_1859_01 darwin_1860_01
will
darwin_1859_01 darwin_1860_01 darwin_1872_01
breed
darwin_1859_01
most
darwin_1859_01 darwin_1860_01 darwin_1872_01
freely under the most unnatural conditions (for instance,
darwin_1859_01 darwin_1860_01
the
darwin_1859_01 darwin_1860_01 darwin_1872_01
rabbit
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
and ferret
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
kept in hutches), showing that their reproductive
darwin_1872_01
organ
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1859_01 darwin_1860_01
ystem
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
h
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
s
darwin_1872_01
re
darwin_1859_01 darwin_1860_01 darwin_1872_01
not
darwin_1859_01 darwin_1860_01
b
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
en thu
darwin_1872_01
a
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1872_01
ily
darwin_1859_01 darwin_1860_01 darwin_1872_01
affected; so will some animals and plants withstand domestication or cultivation, and vary very slightly—perhaps hardly more than in a state of nature.
darwin_1859_01 darwin_1860_01
A
darwin_1872_01
Some naturalists have maintained that all variations are connected with the act of sexual reproduction; but this is certainly an error; for I have given in another work a
darwin_1859_01 darwin_1860_01 darwin_1872_01
long list
darwin_1859_01 darwin_1860_01
could easily be given
darwin_1859_01 darwin_1860_01 darwin_1872_01
of "sporting plants;"
darwin_1872_01
as they are called
darwin_1859_01 darwin_1860_01 darwin_1872_01
by
darwin_1859_01 darwin_1860_01
this term
darwin_1859_01 darwin_1860_01 darwin_1872_01
gardeners
darwin_1872_01
;—that
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
me
darwin_1872_01
is, of pl
darwin_1859_01 darwin_1860_01 darwin_1872_01
an
darwin_1872_01
ts which have suddenly produced
darwin_1859_01 darwin_1860_01 darwin_1872_01
a single bud
darwin_1859_01 darwin_1860_01
or offset,
darwin_1859_01 darwin_1860_01 darwin_1872_01
w
darwin_1859_01 darwin_1860_01
h
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
c
darwin_1872_01
t
darwin_1859_01 darwin_1860_01 darwin_1872_01
h
darwin_1859_01 darwin_1860_01
suddenly assumes
darwin_1859_01 darwin_1860_01 darwin_1872_01
a new and sometimes
darwin_1859_01 darwin_1860_01
v
darwin_1872_01
wid
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
r
darwin_1872_01
l
darwin_1859_01 darwin_1860_01 darwin_1872_01
y different character from that of the
darwin_1872_01
othe
darwin_1859_01 darwin_1860_01 darwin_1872_01
r
darwin_1859_01 darwin_1860_01
e
darwin_1872_01
bud
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1859_01 darwin_1860_01
t
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
f
darwin_1872_01
n
darwin_1859_01 darwin_1860_01 darwin_1872_01
the
darwin_1872_01
same
darwin_1859_01 darwin_1860_01 darwin_1872_01
plant.
darwin_1859_01 darwin_1860_01
Suc
darwin_1872_01
T
darwin_1859_01 darwin_1860_01 darwin_1872_01
h
darwin_1872_01
ese
darwin_1859_01 darwin_1860_01 darwin_1872_01
bud
darwin_1872_01
variation
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1872_01
, as they may be named,
darwin_1859_01 darwin_1860_01 darwin_1872_01
can be propagated by graft
darwin_1859_01 darwin_1860_01
ing
darwin_1872_01
s, offsets
darwin_1859_01 darwin_1860_01 darwin_1872_01
, &c., and sometimes by seed. The
darwin_1859_01 darwin_1860_01
se "sports" are extremel
darwin_1859_01 darwin_1860_01 darwin_1872_01
y
darwin_1872_01
occur
darwin_1859_01 darwin_1860_01 darwin_1872_01
rare
darwin_1872_01
ly
darwin_1859_01 darwin_1860_01 darwin_1872_01
under nature, but
darwin_1872_01
are
darwin_1859_01 darwin_1860_01 darwin_1872_01
far from rare under cult
darwin_1859_01 darwin_1860_01
ivation; and in this case we see that the t
darwin_1872_01
u
darwin_1859_01 darwin_1860_01 darwin_1872_01
re
darwin_1859_01 darwin_1860_01
atment
darwin_1872_01
.
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
of the parent ha
darwin_1872_01
A
darwin_1859_01 darwin_1860_01 darwin_1872_01
s a
darwin_1859_01 darwin_1860_01
ffected
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
a
darwin_1872_01
single
darwin_1859_01 darwin_1860_01 darwin_1872_01
bud o
darwin_1859_01 darwin_1860_01
r offset, and not the ov
darwin_1859_01 darwin_1860_01 darwin_1872_01
u
darwin_1859_01 darwin_1860_01
les or pollen. But it is the opinion of most physiologists that there is no essential difference between a bud and an ovule in their earliest stages of formation; so that, in fact, "sports" support my view, that variability may be largely attributed to the ovules or pollen, or to both, having been affected by the treatmen
darwin_1859_01 darwin_1860_01 darwin_1872_01
t of
darwin_1859_01 darwin_1860_01
the parent prior to the act of conception. These cases
darwin_1872_01
m
darwin_1859_01 darwin_1860_01 darwin_1872_01
any
darwin_1859_01 darwin_1860_01
how show
darwin_1859_01 darwin_1860_01 darwin_1872_01
th
darwin_1872_01
ous
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
t
darwin_1872_01
nds,
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
v
darwin_1872_01
produced ye
darwin_1859_01 darwin_1860_01 darwin_1872_01
ar
darwin_1859_01 darwin_1860_01
i
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1872_01
f
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
i
darwin_1872_01
er year
darwin_1859_01 darwin_1860_01 darwin_1872_01
on
darwin_1859_01 darwin_1860_01
is not necessarily connected, as some authors have supposed, with the act of generation. Seedlings from
darwin_1859_01 darwin_1860_01 darwin_1872_01
the same
darwin_1859_01 darwin_1860_01
frui
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
, and the young of the same litter, sometimes differ considerably from each other, though both the young and the pa
darwin_1859_01 darwin_1860_01 darwin_1872_01
re
darwin_1859_01 darwin_1860_01
nts,
darwin_1872_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
as Müll
darwin_1872_01
und
darwin_1859_01 darwin_1860_01 darwin_1872_01
er
darwin_1859_01 darwin_1860_01
has remarked, have appare
darwin_1872_01
u
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1859_01 darwin_1860_01
tly been exp
darwin_1872_01
if
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
sed to exactly the sa
darwin_1872_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
m
darwin_1859_01 darwin_1860_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
conditions
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1872_01
has been kn
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
f
darwin_1872_01
wn
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1872_01
sudden
darwin_1859_01 darwin_1860_01 darwin_1872_01
l
darwin_1859_01 darwin_1860_01
if
darwin_1872_01
y to assum
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
a new character
darwin_1859_01 darwin_1860_01 darwin_1872_01
; and
darwin_1859_01 darwin_1860_01
th
darwin_1872_01
as buds on d
darwin_1859_01 darwin_1860_01 darwin_1872_01
is
darwin_1872_01
tinct
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1872_01
tree
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1859_01 darwin_1860_01
h
darwin_1872_01
, gr
darwin_1859_01 darwin_1860_01 darwin_1872_01
ow
darwin_1859_01 darwin_1860_01
s how
darwin_1872_01
ing
darwin_1859_01 darwin_1860_01 darwin_1872_01
un
darwin_1872_01
der different conditions, have somet
darwin_1859_01 darwin_1860_01 darwin_1872_01
im
darwin_1859_01 darwin_1860_01
p
darwin_1872_01
es yielded nearly the same variety—f
darwin_1859_01 darwin_1860_01 darwin_1872_01
or
darwin_1872_01
ins
darwin_1859_01 darwin_1860_01 darwin_1872_01
tan
darwin_1872_01
ce, buds on peach-trees producing nectarines, and buds on common roses producing moss-roses—we clearly see tha
darwin_1859_01 darwin_1860_01 darwin_1872_01
t the
darwin_1859_01 darwin_1860_01
di
darwin_1872_01
natu
darwin_1859_01 darwin_1860_01 darwin_1872_01
re
darwin_1859_01 darwin_1860_01
ct effects
darwin_1859_01 darwin_1860_01 darwin_1872_01
of the conditions
darwin_1872_01
is
darwin_1859_01 darwin_1860_01 darwin_1872_01
of
darwin_1859_01 darwin_1860_01
l
darwin_1872_01
subord
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
f
darwin_1872_01
nat
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
import
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
r
darwin_1872_01
nc
darwin_1859_01 darwin_1860_01 darwin_1872_01
e in comparison with the
darwin_1859_01 darwin_1860_01
l
darwin_1872_01
n
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
ws of
darwin_1872_01
tu
darwin_1859_01 darwin_1860_01 darwin_1872_01
re
darwin_1859_01 darwin_1860_01
production,
darwin_1859_01
and
darwin_1859_01 darwin_1860_01
of growth, and of inheritance; for had the action
darwin_1859_01 darwin_1860_01 darwin_1872_01
of the
darwin_1859_01 darwin_1860_01
c
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
nditions been di
darwin_1859_01 darwin_1860_01 darwin_1872_01
r
darwin_1859_01 darwin_1860_01
ect, if
darwin_1872_01
g
darwin_1859_01 darwin_1860_01 darwin_1872_01
an
darwin_1859_01 darwin_1860_01
y of the young had var
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
ed, all would probably have varied
darwin_1872_01
sm
darwin_1859_01 darwin_1860_01 darwin_1872_01
in
darwin_1859_01 darwin_1860_01
the same manner. To judge how much, in the case of any variation, we should attribute to the direct action of heat, moisture, light, food, &c., is most difficult: my impression is, that with animals such agencies have produced very little direct effect, though apparently more in the case of plants. Un
darwin_1859_01 darwin_1860_01 darwin_1872_01
de
darwin_1859_01 darwin_1860_01
r this point of view, Mr. Buckman's recent experiments on plants
darwin_1859_01
s
darwin_1860_01
ar
darwin_1859_01 darwin_1860_01
e
darwin_1859_01
em
darwin_1859_01 darwin_1860_01
extremely valuable. When all or nearly all the individuals exposed to certain conditions are affec
darwin_1859_01 darwin_1860_01 darwin_1872_01
te
darwin_1859_01 darwin_1860_01
d in the same way, the change at fi
darwin_1859_01 darwin_1860_01 darwin_1872_01
r
darwin_1859_01 darwin_1860_01
st appears to be directly due to such conditions; but in some cases it can be shown that quite opposite conditions produce si
darwin_1859_01 darwin_1860_01 darwin_1872_01
mi
darwin_1859_01 darwin_1860_01
l
darwin_1872_01
ning e
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
ch
darwin_1859_01 darwin_1860_01
anges of structure. Nevertheless some slight amount of change may, I think, be attributed to the direct action of the conditions of life—as, in some cases, increased size from amount of food, colour from
darwin_1859_01 darwin_1860_01 darwin_1872_01
particular
darwin_1859_01 darwin_1860_01
kinds
darwin_1872_01
form
darwin_1859_01 darwin_1860_01 darwin_1872_01
of
darwin_1859_01 darwin_1860_01
food
darwin_1859_01
and
darwin_1860_01
or
darwin_1859_01 darwin_1860_01
from light,
darwin_1872_01
v
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1872_01
riatio
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1859_01 darwin_1860_01
d
darwin_1872_01
;—
darwin_1859_01 darwin_1860_01 darwin_1872_01
perhaps
darwin_1872_01
of not more importance than
darwin_1859_01 darwin_1860_01 darwin_1872_01
the
darwin_1872_01
na
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1872_01
ure of the spark, by w
darwin_1859_01 darwin_1860_01 darwin_1872_01
hic
darwin_1859_01 darwin_1860_01
kne
darwin_1872_01
h a ma
darwin_1859_01 darwin_1860_01 darwin_1872_01
ss of
darwin_1859_01 darwin_1860_01
f
darwin_1872_01
combustible matter is ignited, has in determining the nat
darwin_1859_01 darwin_1860_01 darwin_1872_01
ur
darwin_1872_01
e of the
darwin_1859_01 darwin_1860_01 darwin_1872_01
f
darwin_1859_01 darwin_1860_01
r
darwin_1872_01
lames. Effects
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
m climate.
darwin_1872_01
f
darwin_1859_01 darwin_1860_01 darwin_1872_01
Habit a
darwin_1872_01
nd of the Use or Disuse of Parts; Corre
darwin_1859_01 darwin_1860_01 darwin_1872_01
l
darwin_1859_01 darwin_1860_01
s
darwin_1872_01
ated Variati
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1872_01
n; Inheritance. Changed
darwin_1859_01 darwin_1860_01 darwin_1872_01
ha
darwin_1872_01
bit
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1872_01
produce
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1872_01
n
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
d
darwin_1872_01
inh
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
c
darwin_1872_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
d
darwin_1872_01
t
darwin_1859_01 darwin_1860_01 darwin_1872_01
ed
darwin_1859_01 darwin_1860_01
influ
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
n
darwin_1872_01
ffe
darwin_1859_01 darwin_1860_01 darwin_1872_01
c
darwin_1859_01 darwin_1860_01
e
darwin_1872_01
t
darwin_1859_01 darwin_1860_01 darwin_1872_01
, as in the period of
darwin_1872_01
the
darwin_1859_01 darwin_1860_01 darwin_1872_01
flowering
darwin_1859_01 darwin_1860_01
with
darwin_1872_01
of
darwin_1859_01 darwin_1860_01 darwin_1872_01
plants when transported from one climate to another.
darwin_1859_01 darwin_1860_01
In
darwin_1872_01
With
darwin_1859_01 darwin_1860_01 darwin_1872_01
animals
darwin_1872_01
the
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1872_01
ncreased use or disuse of par
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
has
darwin_1872_01
had
darwin_1859_01 darwin_1860_01 darwin_1872_01
a more marked
darwin_1859_01 darwin_1860_01
effect; for
darwin_1859_01 darwin_1860_01 darwin_1872_01
in
darwin_1859_01 darwin_1860_01
sta
darwin_1872_01
flue
darwin_1859_01 darwin_1860_01 darwin_1872_01
nce
darwin_1859_01 darwin_1860_01
,
darwin_1872_01
; thus
darwin_1859_01 darwin_1860_01 darwin_1872_01
I find in the domestic duck that the bones of the wing weigh less and the bones of the leg more, in proportion to the whole skeleton, than do the same bones in the wild
darwin_1859_01 darwin_1860_01
-
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
duck; and
darwin_1859_01 darwin_1860_01
I presume that
darwin_1859_01 darwin_1860_01 darwin_1872_01
this change may be safely attributed to the domestic duck flying much less, and walking more, than its wild parent
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
. The great and inherited development of the udders in cows and goats in countries where they are habitually milked, in comparison with the
darwin_1859_01 darwin_1860_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1859_01 darwin_1860_01
tate of thes
darwin_1859_01 darwin_1860_01 darwin_1872_01
e organs in other countries, is
darwin_1872_01
probably
darwin_1859_01 darwin_1860_01 darwin_1872_01
another instance of the effect
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
of use. Not
darwin_1859_01 darwin_1860_01
a
darwin_1872_01
one
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
single
darwin_1872_01
of our
darwin_1859_01 darwin_1860_01 darwin_1872_01
domestic animal
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
can be named which has not in some country drooping ears; and the view
darwin_1872_01
which has been
darwin_1859_01 darwin_1860_01 darwin_1872_01
suggested
darwin_1859_01 darwin_1860_01
by some authors,
darwin_1859_01 darwin_1860_01 darwin_1872_01
that the drooping is due to the disuse of the muscles of the ear, from the animals
darwin_1859_01 darwin_1860_01
not
darwin_1859_01 darwin_1860_01 darwin_1872_01
being
darwin_1872_01
seldom
darwin_1859_01 darwin_1860_01 darwin_1872_01
much alarmed
darwin_1859_01 darwin_1860_01
by danger
darwin_1859_01 darwin_1860_01 darwin_1872_01
, seems probable.
darwin_1859_01 darwin_1860_01
There are m
darwin_1872_01
M
darwin_1859_01 darwin_1860_01 darwin_1872_01
any laws regulat
darwin_1859_01 darwin_1860_01
ing
darwin_1872_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
variation, some few of which can be dimly seen, and will
darwin_1859_01 darwin_1860_01
be
darwin_1859_01 darwin_1860_01 darwin_1872_01
hereafter b
darwin_1872_01
e b
darwin_1859_01 darwin_1860_01 darwin_1872_01
riefly
darwin_1859_01 darwin_1860_01
ment
darwin_1872_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
on
darwin_1872_01
scuss
darwin_1859_01 darwin_1860_01 darwin_1872_01
ed. I will here only allude to what may be called correlat
darwin_1872_01
ed variat
darwin_1859_01 darwin_1860_01 darwin_1872_01
ion
darwin_1859_01 darwin_1860_01
of growth
darwin_1859_01 darwin_1860_01 darwin_1872_01
.
darwin_1859_01 darwin_1860_01
A
darwin_1872_01
Importa
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1859_01 darwin_1860_01
y
darwin_1872_01
t
darwin_1859_01 darwin_1860_01 darwin_1872_01
change
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
in the embryo or larva will
darwin_1859_01 darwin_1860_01
almost ce
darwin_1872_01
p
darwin_1859_01 darwin_1860_01 darwin_1872_01
r
darwin_1859_01 darwin_1860_01
t
darwin_1872_01
ob
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
in
darwin_1872_01
b
darwin_1859_01 darwin_1860_01 darwin_1872_01
ly entail changes in the mature animal. In monstrosities, the correlations between quite distinct parts are very curious; and many instances are given in Isidore Geoffroy St. Hilaire's great work on this subject. Breeders believe that long limbs are almost always accompanied by an elongated head. Some instances of correlation are quite whimsical
darwin_1859_01 darwin_1860_01
:
darwin_1872_01
;
darwin_1859_01 darwin_1860_01 darwin_1872_01
thus cats w
darwin_1872_01
hich are entirely wh
darwin_1859_01 darwin_1860_01 darwin_1872_01
it
darwin_1872_01
e and
darwin_1859_01 darwin_1860_01 darwin_1872_01
h
darwin_1872_01
ave
darwin_1859_01 darwin_1860_01 darwin_1872_01
blue eyes are
darwin_1859_01 darwin_1860_01
i
darwin_1872_01
ge
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1859_01 darwin_1860_01
va
darwin_1872_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
r
darwin_1859_01 darwin_1860_01
i
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
b
darwin_1872_01
l
darwin_1859_01 darwin_1860_01 darwin_1872_01
ly deaf;
darwin_1872_01
but it has been lately stated by Mr. Tait that this is
darwin_1859_01 darwin_1860_01 darwin_1872_01
c
darwin_1872_01
onfined to the males. C
darwin_1859_01 darwin_1860_01 darwin_1872_01
olour and constitutional peculiarities go together, of which many remarkable cases could be given amongst animals and plants. From
darwin_1859_01 darwin_1860_01
the
darwin_1859_01 darwin_1860_01 darwin_1872_01
facts collected by Heusinger, it appears that white sheep and pigs are
darwin_1859_01 darwin_1860_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
ffere
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1859_01 darwin_1860_01
tly affect
darwin_1872_01
jur
darwin_1859_01 darwin_1860_01 darwin_1872_01
ed
darwin_1859_01 darwin_1860_01
f
darwin_1872_01
by ce
darwin_1859_01 darwin_1860_01 darwin_1872_01
r
darwin_1859_01 darwin_1860_01
om
darwin_1872_01
tain
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1872_01
plants, whilst dark-
darwin_1859_01 darwin_1860_01 darwin_1872_01
coloured individuals
darwin_1859_01 darwin_1860_01
b
darwin_1872_01
escape: Professor Wyman has recentl
darwin_1859_01 darwin_1860_01 darwin_1872_01
y c
darwin_1872_01
ommunicated to me a good illustration of this fact; on asking some farm
darwin_1859_01 darwin_1860_01 darwin_1872_01
er
darwin_1872_01
s in Virginia how i
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1872_01
was that all their pigs were black, they informed him that the pigs ate the p
darwin_1859_01 darwin_1860_01 darwin_1872_01
ain
darwin_1872_01
t-root (Lachnanthes), which coloured their bones pink, and which caused the hoofs of all but the black
darwin_1859_01 darwin_1860_01 darwin_1872_01
v
darwin_1859_01 darwin_1860_01
eg
darwin_1872_01
ari
darwin_1859_01 darwin_1860_01 darwin_1872_01
et
darwin_1872_01
ies to drop off;
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1872_01
nd one of the "crackers" (i.e. Virginia squatters) added, "we select the
darwin_1859_01 darwin_1860_01 darwin_1872_01
bl
darwin_1872_01
ack members of a litter for raising, as they alon
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
p
darwin_1872_01
have a g
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1872_01
od chance of l
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
so
darwin_1872_01
vi
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1859_01 darwin_1860_01
s
darwin_1872_01
g
darwin_1859_01 darwin_1860_01 darwin_1872_01
.
darwin_1872_01
"
darwin_1859_01 darwin_1860_01 darwin_1872_01
Hairless dogs have imperfect teeth
darwin_1860_01 darwin_1872_01
:
darwin_1859_01
;
darwin_1859_01 darwin_1860_01 darwin_1872_01
long-haired and coarse-haired animals are apt to have, as is asserted, long or many horns; pigeons with feathered feet have skin between their outer toes; pigeons with short beaks have small feet, and those with long beaks large feet. Hence
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
if man goes on selecting, and thus augmenting, any peculiarity, he will almost certainly
darwin_1859_01 darwin_1860_01
unconsciously
darwin_1859_01 darwin_1860_01 darwin_1872_01
modify
darwin_1872_01
unintentionally
darwin_1859_01 darwin_1860_01 darwin_1872_01
other parts of the structure, owing to the mysterious laws of
darwin_1859_01 darwin_1860_01
the
darwin_1859_01 darwin_1860_01 darwin_1872_01
correlation
darwin_1859_01 darwin_1860_01
of growth
darwin_1859_01 darwin_1860_01 darwin_1872_01
. The result
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
of the various,
darwin_1859_01 darwin_1860_01
quite
darwin_1859_01 darwin_1860_01 darwin_1872_01
unknown, or
darwin_1872_01
but
darwin_1859_01 darwin_1860_01 darwin_1872_01
dimly
darwin_1872_01
under
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1859_01 darwin_1860_01
een
darwin_1872_01
tood
darwin_1859_01 darwin_1860_01 darwin_1872_01
laws of variation
darwin_1859_01 darwin_1860_01
is
darwin_1872_01
are
darwin_1859_01 darwin_1860_01 darwin_1872_01
infinitely complex and diversified. It is well worth while carefully to study the several treatises
darwin_1859_01 darwin_1860_01
published
darwin_1859_01 darwin_1860_01 darwin_1872_01
on some of our old cultivated plants, as on the hyacinth, potato, even the dahlia, &c.; and it is really surprising to note the endless points
darwin_1859_01 darwin_1860_01
in
darwin_1872_01
of
darwin_1859_01 darwin_1860_01 darwin_1872_01
structure and constitution in which the varieties and sub-varieties differ slightly from each other. The whole organisation seems to have become plastic, and
darwin_1859_01 darwin_1860_01
tends to
darwin_1859_01 darwin_1860_01 darwin_1872_01
depart
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
in
darwin_1859_01 darwin_1860_01
some
darwin_1872_01
a
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1859_01 darwin_1860_01
ma
darwin_1859_01 darwin_1860_01 darwin_1872_01
l
darwin_1859_01 darwin_1860_01
l
darwin_1872_01
ight
darwin_1859_01 darwin_1860_01 darwin_1872_01
degree from that of the parental type. Any variation which is not inherited is unimportant for us. But the number and diversity of inheritable deviations of structure, both those of slight and those of considerable physiological importance,
darwin_1859_01 darwin_1860_01
is
darwin_1872_01
are
darwin_1859_01 darwin_1860_01 darwin_1872_01
endless. Dr. Prosper Lucas's treatise, in two large volumes, is the fullest and the best on this subject. No breeder doubts how strong is the tendency to inheritance
darwin_1859_01 darwin_1860_01
:
darwin_1872_01
; that
darwin_1859_01 darwin_1860_01 darwin_1872_01
like produces like is his fundamental belief: doubts have been thrown on this principle
darwin_1872_01
only
darwin_1859_01 darwin_1860_01 darwin_1872_01
by theoretical writers
darwin_1859_01 darwin_1860_01
alone
darwin_1859_01 darwin_1860_01 darwin_1872_01
. When a
darwin_1860_01 darwin_1872_01
ny
darwin_1859_01 darwin_1860_01 darwin_1872_01
deviation
darwin_1860_01 darwin_1872_01
of structure often
darwin_1859_01 darwin_1860_01 darwin_1872_01
appears
darwin_1859_01
not unfrequently
darwin_1859_01 darwin_1860_01 darwin_1872_01
, and we see it in the father and child, we cannot tell whether it may not be due to the same
darwin_1859_01
original
darwin_1859_01 darwin_1860_01 darwin_1872_01
cause
darwin_1860_01 darwin_1872_01
having
darwin_1859_01 darwin_1860_01 darwin_1872_01
act
darwin_1860_01 darwin_1872_01
ed
darwin_1859_01
ing
darwin_1859_01 darwin_1860_01 darwin_1872_01
on both; but when amongst individuals, apparently exposed to the same conditions, any very rare deviation, due to some extraordinary combination of circumstances, appears in the parent—say, once among
darwin_1859_01 darwin_1860_01
st
darwin_1859_01 darwin_1860_01 darwin_1872_01
several million individuals
darwin_1860_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
—and it reappears in the child, the mere doctrine of chances almost compels us to attribute its reappearance to inheritance. Every one must have heard of cases of albinism, prickly skin, hairy bodies, &c., appearing in several members of the same family. If strange and rare deviations of structure are
darwin_1859_01 darwin_1860_01
t
darwin_1859_01 darwin_1860_01 darwin_1872_01
r
darwin_1859_01 darwin_1860_01
u
darwin_1872_01
eal
darwin_1859_01 darwin_1860_01 darwin_1872_01
ly inherited, less strange and commoner deviations may be freely admitted to be inheritable. Perhaps the correct way of viewing the whole subject
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
would be, to look at the inheritance of every character whatever as the rule, and non-inheritance as the anomaly. The laws governing inheritance are
darwin_1859_01 darwin_1860_01
qui
darwin_1872_01
for
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1872_01
h
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
most part
darwin_1859_01 darwin_1860_01 darwin_1872_01
unknown
darwin_1859_01 darwin_1860_01
;
darwin_1872_01
.
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
n
darwin_1872_01
N
darwin_1859_01 darwin_1860_01 darwin_1872_01
o one can say why
darwin_1859_01 darwin_1872_01
the s
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1872_01
me
darwin_1859_01 darwin_1860_01 darwin_1872_01
peculiarity in different individuals of the same species,
darwin_1860_01 darwin_1872_01
or
darwin_1859_01
and
darwin_1859_01 darwin_1860_01 darwin_1872_01
in
darwin_1859_01 darwin_1860_01
individuals of
darwin_1859_01 darwin_1860_01 darwin_1872_01
different species, is sometimes inherited and sometimes not so; why the child often reverts in certain character
darwin_1872_01
i
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1872_01
tics
darwin_1859_01 darwin_1860_01 darwin_1872_01
to its grandfather or grandmother or
darwin_1859_01 darwin_1860_01
other
darwin_1859_01 darwin_1860_01 darwin_1872_01
m
darwin_1859_01
uch m
darwin_1859_01 darwin_1860_01 darwin_1872_01
ore remote ancestor; why a peculiarity is often transmitted from one sex to both sexes, or to one sex alone, more commonly but not exclusively to the like sex. It is a fact of some
darwin_1859_01 darwin_1860_01
little
darwin_1859_01 darwin_1860_01 darwin_1872_01
importance to us, that peculiarities appearing in the males of our domestic breeds are often transmitted
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
either exclusively
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
or in a much greater degree, to
darwin_1872_01
the
darwin_1859_01 darwin_1860_01 darwin_1872_01
males alone. A much more important rule, which I think may be trusted, is that, at whatever period of life a peculiarity first appears, it tends to
darwin_1872_01
re-
darwin_1859_01 darwin_1860_01 darwin_1872_01
appear in the offspring at a corresponding age, though sometimes earlier. In many cases this could not be otherwise
darwin_1859_01 darwin_1860_01
:
darwin_1872_01
;
darwin_1859_01 darwin_1860_01 darwin_1872_01
thus the inherited peculiarities in the horns of cattle could appear only in the offspring when nearly mature; peculiarities in the silk
darwin_1872_01
-
darwin_1859_01 darwin_1860_01 darwin_1872_01
worm are known to appear at the corresponding caterpillar or cocoon stage. But hereditary diseases and some other facts make me believe that the rule has a wider extension, and that
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
when there is no apparent reason why a peculiarity should appear at any particular age, yet that it does tend to appear in the offspring at the same period at which it first appeared in the parent. I believe this rule to be of the highest importance in explaining the laws of embryology. These remarks are of course confined to the first appearance of the peculiarity, and not to
darwin_1859_01 darwin_1860_01
i
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
s
darwin_1872_01
he
darwin_1859_01 darwin_1860_01 darwin_1872_01
primary cause
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
which may have acted on the ovules or
darwin_1872_01
on the
darwin_1859_01 darwin_1860_01 darwin_1872_01
male element; in nearly the same manner as
darwin_1872_01
the increased length of the horns
darwin_1859_01 darwin_1860_01 darwin_1872_01
in the
darwin_1859_01 darwin_1860_01
crossed
darwin_1859_01 darwin_1860_01 darwin_1872_01
offspring from a short-horned cow by a long-horned bull
darwin_1859_01 darwin_1860_01
, the greater length of horn
darwin_1859_01 darwin_1860_01 darwin_1872_01
, though appearing late in life, is clearly due to the male element. Having alluded to the subject of reversion, I may here refer to a statement often made by naturalists—namely, that our domestic varieties, when run wild, gradually but
darwin_1859_01 darwin_1860_01
certa
darwin_1859_01 darwin_1860_01 darwin_1872_01
in
darwin_1872_01
variab
darwin_1859_01 darwin_1860_01 darwin_1872_01
ly revert in character to their aboriginal stocks. Hence it has been argued that no deductions can be drawn from domestic races to species in a state of nature. I have in vain endeavoured to discover on what decisive facts the above statement has so often and so boldly been made. There would be great difficulty in proving its truth: we may safely conclude that very many of the most strongly
darwin_1859_01 darwin_1860_01
-
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
marked domestic varieties could not possibly live in a wild state. In many cases we do not know what the aboriginal stock was, and so could not tell whether or not nearly perfect reversion had ensued. It would be
darwin_1859_01 darwin_1860_01
quite
darwin_1859_01 darwin_1860_01 darwin_1872_01
necessary
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
in order to prevent the effects of intercrossing, that only a single variety should
darwin_1872_01
have
darwin_1859_01 darwin_1860_01 darwin_1872_01
be
darwin_1872_01
en
darwin_1859_01 darwin_1860_01 darwin_1872_01
turned loose in its new home. Nevertheless, as our varieties certainly do occasionally revert in some of their characters to ancestral forms, it seems to me not improbable
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
that if we could succeed in naturalising, or were to cultivate, during many generations, the several races, for instance, of the cabbage, in very poor soil (in which case, however, some effect would have to be attributed to the d
darwin_1872_01
ef
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
rec
darwin_1872_01
ni
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1872_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
action of the poor soil), that they would
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
to a large extent, or even wholly, revert to the wild aboriginal stock. Whether or not the experiment would succeed, is not of great importance for our line of argument; for by the experiment itself the conditions of life are changed. If it could be shown that our domestic varieties manifested a strong tendency to reversion,—that is, to lose their acquired characters, whilst kept under
darwin_1860_01 darwin_1872_01
t
darwin_1859_01
unc
darwin_1859_01 darwin_1860_01 darwin_1872_01
h
darwin_1860_01 darwin_1872_01
e s
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1860_01 darwin_1872_01
m
darwin_1859_01
ng
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
conditions, and whilst kept in a considerable body, so that free intercrossing might check, by blending together, any slight deviations
darwin_1860_01 darwin_1872_01
in their
darwin_1859_01
of
darwin_1859_01 darwin_1860_01 darwin_1872_01
structure, in such case, I grant that we could deduce nothing from domestic varieties in regard to species. But there is not a shadow of evidence in favour of this view: to assert that we could not breed our cart and race-horses, long and short-horned cattle, and poultry of various breeds, and esculent vegetables, for an
darwin_1859_01 darwin_1860_01
almost i
darwin_1872_01
u
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1859_01 darwin_1860_01
f
darwin_1872_01
l
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
n
darwin_1872_01
m
darwin_1859_01 darwin_1860_01 darwin_1872_01
ite
darwin_1872_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
number of generations, would be opposed to all experience.
darwin_1859_01 darwin_1860_01
I may add, that when under nature the conditions of life do change, variations and reversions of c
darwin_1872_01
C
darwin_1859_01 darwin_1860_01 darwin_1872_01
haracter
darwin_1859_01 darwin_1860_01
pr
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
bably
darwin_1872_01
f
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
d
darwin_1872_01
D
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
occur; but natural s
darwin_1872_01
m
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
lec
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
ti
darwin_1859_01 darwin_1860_01
on, as will hereafter be explained, will determine how far the new
darwin_1859_01 darwin_1860_01 darwin_1872_01
c
darwin_1859_01 darwin_1860_01
haracters
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
thus
darwin_1872_01
V
darwin_1859_01 darwin_1860_01 darwin_1872_01
ari
darwin_1872_01
etie
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1872_01
: difficulty of dist
darwin_1859_01 darwin_1860_01 darwin_1872_01
ing
darwin_1859_01 darwin_1860_01
darwin_1872_01
ui
darwin_1859_01 darwin_1860_01 darwin_1872_01
sh
darwin_1859_01 darwin_1860_01
all
darwin_1872_01
ing
darwin_1859_01 darwin_1860_01 darwin_1872_01
be
darwin_1872_01
tween
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1872_01
Varieties and S
darwin_1859_01 darwin_1860_01 darwin_1872_01
p
darwin_1872_01
ecies; origin of Domestic Varieties from one or mo
darwin_1859_01 darwin_1860_01 darwin_1872_01
re
darwin_1872_01
Specie
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1859_01 darwin_1860_01
erved
darwin_1859_01 darwin_1860_01 darwin_1872_01
. When we look to the hereditary varieties or races of our domestic animals and plants, and compare them with
darwin_1859_01
species
darwin_1859_01 darwin_1860_01 darwin_1872_01
closely allied
darwin_1860_01 darwin_1872_01
sp
darwin_1859_01
tog
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1860_01 darwin_1872_01
ci
darwin_1859_01
th
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1860_01 darwin_1872_01
s
darwin_1859_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
, we generally perceive in each domestic race, as already remarked, less uniformity of character than in true species. Domestic races of
darwin_1859_01 darwin_1860_01
the same species, also, of
darwin_1859_01 darwin_1860_01 darwin_1872_01
ten have a somewhat monstrous character; by which I mean, that, although differing from each other, and from
darwin_1859_01
the
darwin_1859_01 darwin_1860_01 darwin_1872_01
other species of the same genus, in several trifling respects, they often differ in an extreme degree in some one part, both when compared one with another, and more especially when compared with
darwin_1859_01 darwin_1860_01
all
darwin_1859_01 darwin_1860_01 darwin_1872_01
the species
darwin_1859_01 darwin_1860_01
i
darwin_1872_01
u
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1872_01
der
darwin_1859_01 darwin_1860_01 darwin_1872_01
nature to which they are nearest allied. With these exceptions (and with that of the perfect fertility of varieties when crossed,—a subject hereafter to be discussed), domestic races of the same species differ from each other in the same manner as
darwin_1859_01 darwin_1860_01
, only in most cases in a lesser degree than,
darwin_1859_01 darwin_1860_01 darwin_1872_01
do
darwin_1872_01
the
darwin_1859_01 darwin_1860_01 darwin_1872_01
closely
darwin_1859_01 darwin_1860_01
-
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
allied species of the same genus in a state of nature
darwin_1859_01 darwin_1860_01
.
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
I
darwin_1872_01
but
darwin_1859_01 darwin_1860_01 darwin_1872_01
th
darwin_1872_01
e differences
darwin_1859_01 darwin_1860_01 darwin_1872_01
in
darwin_1859_01 darwin_1860_01
k
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1872_01
mos
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1872_01
cases are less in degree. T
darwin_1859_01 darwin_1860_01 darwin_1872_01
his must be admitted
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
when we find
darwin_1872_01
as
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
hat
darwin_1872_01
rue, for
darwin_1859_01 darwin_1860_01 darwin_1872_01
the
darwin_1859_01 darwin_1860_01
re are hardly any
darwin_1859_01 darwin_1860_01 darwin_1872_01
domestic races
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
either am
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1872_01
f ma
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1859_01 darwin_1860_01
gst
darwin_1872_01
y
darwin_1859_01 darwin_1860_01 darwin_1872_01
animals
darwin_1859_01 darwin_1860_01
or
darwin_1872_01
and
darwin_1859_01 darwin_1860_01 darwin_1872_01
plants
darwin_1859_01 darwin_1860_01
, which
darwin_1859_01 darwin_1860_01 darwin_1872_01
have
darwin_1859_01 darwin_1860_01
not
darwin_1859_01 darwin_1860_01 darwin_1872_01
been ranked by
darwin_1859_01
some
darwin_1859_01 darwin_1860_01
c
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
om
darwin_1859_01 darwin_1860_01
p
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
tent judges as mere varieties, and by other
darwin_1859_01 darwin_1860_01 darwin_1872_01
competent judges as the descendants of aboriginally distinct species
darwin_1872_01
, and by other competent judges as mere varieties
darwin_1859_01 darwin_1860_01 darwin_1872_01
. If any
darwin_1872_01
well
darwin_1859_01 darwin_1860_01 darwin_1872_01
marked distinction existed between
darwin_1872_01
a
darwin_1859_01 darwin_1860_01 darwin_1872_01
domestic race
darwin_1859_01 darwin_1860_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
and
darwin_1872_01
a
darwin_1859_01 darwin_1860_01 darwin_1872_01
species, this source of doubt
darwin_1859_01 darwin_1860_01
c
darwin_1872_01
w
darwin_1859_01 darwin_1860_01 darwin_1872_01
ould not so perpetually recur. It has often been stated that domestic races do not differ from each other in characters of generic value. I
darwin_1859_01 darwin_1860_01
think i
darwin_1859_01 darwin_1860_01 darwin_1872_01
t c
darwin_1859_01 darwin_1860_01
ould
darwin_1872_01
an
darwin_1859_01 darwin_1860_01 darwin_1872_01
be shown that this statement is
darwin_1859_01 darwin_1860_01
hardly
darwin_1872_01
not
darwin_1859_01 darwin_1860_01 darwin_1872_01
correct; but naturalists differ
darwin_1859_01
most
darwin_1859_01 darwin_1860_01
widely
darwin_1872_01
much
darwin_1859_01 darwin_1860_01 darwin_1872_01
in determining what characters are of generic value; all such valuations being at present empirical.
darwin_1859_01 darwin_1860_01
Moreover, on t
darwin_1872_01
W
darwin_1859_01 darwin_1860_01 darwin_1872_01
he
darwin_1872_01
n
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
v
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1872_01
t is
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
xplained ho
darwin_1859_01 darwin_1860_01 darwin_1872_01
w
darwin_1859_01 darwin_1860_01
of th
darwin_1872_01
g
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
nera
darwin_1859_01 darwin_1860_01 darwin_1872_01
origin
darwin_1859_01 darwin_1860_01
of g
darwin_1872_01
at
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
u
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1872_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
er
darwin_1872_01
n
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1872_01
ture, it
darwin_1859_01 darwin_1860_01 darwin_1872_01
w
darwin_1859_01 darwin_1860_01
h
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
ch I sha
darwin_1859_01 darwin_1860_01 darwin_1872_01
ll
darwin_1859_01 darwin_1860_01
pr
darwin_1872_01
b
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
se
darwin_1859_01 darwin_1860_01
ntly giv
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
,
darwin_1872_01
n that
darwin_1859_01 darwin_1860_01 darwin_1872_01
we have no right to expect often to
darwin_1859_01 darwin_1860_01
meet w
darwin_1872_01
f
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
th
darwin_1872_01
nd a
darwin_1859_01 darwin_1860_01 darwin_1872_01
generic
darwin_1872_01
amount of
darwin_1859_01 darwin_1860_01 darwin_1872_01
difference
darwin_1859_01 darwin_1860_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
in our domesticated
darwin_1859_01 darwin_1860_01
p
darwin_1859_01 darwin_1860_01 darwin_1872_01
r
darwin_1859_01 darwin_1860_01
odu
darwin_1872_01
a
darwin_1859_01 darwin_1860_01 darwin_1872_01
c
darwin_1859_01 darwin_1860_01
tion
darwin_1872_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
s.
darwin_1859_01 darwin_1860_01
Whe
darwin_1872_01
I
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1859_01 darwin_1860_01
we
darwin_1859_01 darwin_1860_01 darwin_1872_01
attempt
darwin_1872_01
ing
darwin_1859_01 darwin_1860_01 darwin_1872_01
to estimate the amount of structural difference between
darwin_1859_01 darwin_1860_01
th
darwin_1872_01
alli
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
domestic rac
darwin_1859_01 darwin_1860_01
es of the same speci
darwin_1859_01 darwin_1860_01 darwin_1872_01
es, we are soon involved in doubt, from not knowing whether they
darwin_1859_01 darwin_1860_01
h
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
v
darwin_1872_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
e descended from one or several parent
darwin_1859_01
-
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
species. This point, if it could be cleared up, would be interesting; if, for instance, it could be shown that the greyhound, bloodhound, terrier, spaniel, and bull-dog, which we all know propagate their kind
darwin_1859_01 darwin_1860_01
so
darwin_1859_01 darwin_1860_01 darwin_1872_01
truly, were the offspring of any single species, then such facts would have great weight in making us doubt about the immutability of the many
darwin_1859_01 darwin_1860_01
very
darwin_1859_01 darwin_1860_01 darwin_1872_01
closely allied
darwin_1859_01
and
darwin_1859_01 darwin_1860_01 darwin_1872_01
natural species—for instance, of the many foxes—inhabiting different quarters of the world. I do not believe, as we shall presently see, that
darwin_1860_01 darwin_1872_01
the whole amount of difference between the several breeds of the dog has been produced under domestication; I believe that
darwin_1860_01
some
darwin_1872_01
a
darwin_1860_01 darwin_1872_01
sm
darwin_1859_01 darwin_1860_01 darwin_1872_01
all
darwin_1860_01 darwin_1872_01
part
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1860_01 darwin_1872_01
f the difference is d
darwin_1859_01 darwin_1860_01 darwin_1872_01
u
darwin_1860_01 darwin_1872_01
e to thei
darwin_1859_01 darwin_1860_01 darwin_1872_01
r
darwin_1860_01 darwin_1872_01
b
darwin_1859_01
dogs hav
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1860_01 darwin_1872_01
ing
darwin_1859_01 darwin_1860_01 darwin_1872_01
descended from
darwin_1859_01
any one wil
darwin_1859_01 darwin_1860_01 darwin_1872_01
d
darwin_1860_01 darwin_1872_01
istinct
darwin_1859_01 darwin_1860_01 darwin_1872_01
species
darwin_1860_01 darwin_1872_01
.
darwin_1859_01
;
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1860_01 darwin_1872_01
I
darwin_1859_01
but, i
darwin_1859_01 darwin_1860_01 darwin_1872_01
n the case of s
darwin_1872_01
trongly marked races of s
darwin_1859_01 darwin_1860_01 darwin_1872_01
ome other domestic
darwin_1860_01 darwin_1872_01
ated
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1860_01 darwin_1872_01
spe
darwin_1859_01
ra
darwin_1859_01 darwin_1860_01 darwin_1872_01
c
darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01 darwin_1872_01
es, there is presumptive
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
or even strong
darwin_1859_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
evidence
darwin_1860_01 darwin_1872_01
,
darwin_1859_01
in favour of
darwin_1859_01 darwin_1860_01 darwin_1872_01
th
darwin_1860_01 darwin_1872_01
at all
darwin_1860_01
the breeds h
darwin_1860_01 darwin_1872_01
a
darwin_1860_01
v
darwin_1872_01
r
darwin_1860_01 darwin_1872_01
e de
darwin_1859_01
i
darwin_1860_01 darwin_1872_01 darwin_1859_01
s
darwin_1860_01 darwin_1872_01
cended
darwin_1860_01 darwin_1872_01 darwin_1859_01
darwin_1860_01 darwin_1872_01
from a s
darwin_1859_01
v
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1860_01 darwin_1872_01
ngl
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
w
darwin_1860_01 darwin_1872_01
ild stock
darwin_1859_01 darwin_1860_01 darwin_1872_01
. It has often been assumed that man has chosen for domestication animals and plants having an extraordinary inherent tendency to vary, and likewise to withstand diverse climates. I do not dispute that these capacities have added largely to the value of most of our domesticated productions; but how could a savage possibly know, when he first tamed an animal, whether it would vary in succeeding generations, and whether it would endure other climates? Has the little variability of the ass
darwin_1859_01 darwin_1860_01
or
darwin_1872_01
and
darwin_1859_01 darwin_1860_01 darwin_1872_01
g
darwin_1859_01 darwin_1860_01
uinea-f
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
wl
darwin_1872_01
ose
darwin_1859_01 darwin_1860_01 darwin_1872_01
, or the small power of endurance of warmth by the rein
darwin_1859_01
-
darwin_1859_01 darwin_1860_01 darwin_1872_01
deer, or of cold by the common camel, prevented their domestication? I cannot doubt that if other animals and plants, equal in number to our domesticated productions, and belonging to equally diverse classes and countries, were taken from a state of nature, and could be made to breed for an equal number of generations under domestication, they would
darwin_1859_01 darwin_1860_01
vary
darwin_1859_01 darwin_1860_01 darwin_1872_01
on an average
darwin_1872_01
vary
darwin_1859_01 darwin_1860_01 darwin_1872_01
as largely as the parent species of our existing domesticated productions have varied. In the case of most of our anciently domesticated animals and plants,
darwin_1859_01 darwin_1860_01
I do not think
darwin_1859_01 darwin_1860_01 darwin_1872_01
it is
darwin_1872_01
not
darwin_1859_01 darwin_1860_01 darwin_1872_01
possible to come to any definite conclusion, whether they
darwin_1859_01 darwin_1860_01
h
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
v
darwin_1872_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
e descended from one or several
darwin_1860_01 darwin_1872_01
wild
darwin_1859_01 darwin_1860_01 darwin_1872_01
species. The argument mainly relied on by those who believe in the multiple origin of our domestic animals is, that we find in the most ancient
darwin_1859_01 darwin_1860_01
records,
darwin_1872_01
ti
darwin_1859_01 darwin_1860_01 darwin_1872_01
m
darwin_1859_01 darwin_1860_01
ore
darwin_1859_01 darwin_1860_01 darwin_1872_01
es
darwin_1859_01 darwin_1860_01
pecially
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
on the monuments of Egypt,
darwin_1872_01
and in the lake-habitations of Switzerland,
darwin_1859_01 darwin_1860_01 darwin_1872_01
much diversity in the breeds; and that some of the
darwin_1872_01
se ancient
darwin_1859_01 darwin_1860_01 darwin_1872_01
breeds closely resemble,
darwin_1859_01 darwin_1860_01
pe
darwin_1872_01
o
darwin_1859_01 darwin_1860_01 darwin_1872_01
r
darwin_1859_01 darwin_1860_01
haps
darwin_1859_01 darwin_1860_01 darwin_1872_01
are
darwin_1872_01
even
darwin_1859_01 darwin_1860_01 darwin_1872_01
identical with, those still existing.
darwin_1859_01 darwin_1860_01
Even if
darwin_1872_01
But
darwin_1859_01 darwin_1860_01 darwin_1872_01
this
darwin_1872_01
on
darwin_1859_01 darwin_1860_01 darwin_1872_01
l
darwin_1859_01 darwin_1860_01
a
darwin_1872_01
y
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
te
darwin_1872_01
h
darwin_1859_01 darwin_1860_01 darwin_1872_01
r
darwin_1872_01
ows
darwin_1859_01 darwin_1860_01 darwin_1872_01
fa
darwin_1872_01
r ba
darwin_1859_01 darwin_1860_01 darwin_1872_01
c
darwin_1872_01
kwards
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1872_01
he history of civilisation, and shows that animals
darwin_1859_01 darwin_1860_01 darwin_1872_01
were
darwin_1872_01
domesticated at a much earlier period than has hitherto been supposed. The lake-inhabitants o
darwin_1859_01 darwin_1860_01 darwin_1872_01
f
darwin_1859_01 darwin_1860_01
ou
darwin_1872_01
Switzerla
darwin_1859_01 darwin_1860_01 darwin_1872_01
nd
darwin_1859_01 darwin_1860_01
m
darwin_1872_01
cultivated several kinds of wheat and barley, the pea, the poppy f
darwin_1859_01 darwin_1860_01 darwin_1872_01
or
darwin_1872_01
oil, and flax; and they possessed several domesticated animals. They also carried on commerc
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
with other nations. All this clearly shows, as Heer has remarked, that they had at this early age progressed considerably in civilisation; and this again implies a long continued previous period of less advanced civilisation, during which the domesticated animals, kept by different tribes in different di
darwin_1859_01 darwin_1860_01 darwin_1872_01
strict
darwin_1872_01
s, might have varied and given rise to distinct races. Since the discovery of flint tools in the superficial formations of many parts of the world, all geologists believe that barbarian man existed at an enormously remote period; and we know that at the present day there is hard
darwin_1859_01 darwin_1860_01 darwin_1872_01
ly a
darwin_1859_01 darwin_1860_01
nd generally
darwin_1859_01 darwin_1860_01 darwin_1872_01
tr
darwin_1859_01 darwin_1860_01
ue than seems to me to
darwin_1872_01
i
darwin_1859_01 darwin_1860_01 darwin_1872_01
be
darwin_1859_01 darwin_1860_01
the case, what does it show, but that
darwin_1859_01 darwin_1860_01 darwin_1872_01
so
darwin_1859_01 darwin_1860_01
me of our
darwin_1859_01 darwin_1860_01 darwin_1872_01
b
darwin_1859_01 darwin_1860_01
reeds originated there, four or five thousand ye
darwin_1859_01 darwin_1860_01 darwin_1872_01
ar
darwin_1859_01 darwin_1860_01
s ago? But Mr. Horner's researches have rendered it in some degree pro
darwin_1859_01 darwin_1860_01 darwin_1872_01
ba
darwin_1859_01 darwin_1860_01
ble that man s
darwin_1872_01
ro
darwin_1859_01 darwin_1860_01 darwin_1872_01
u
darwin_1859_01 darwin_1860_01
fficiently
darwin_1872_01
s,
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
civilized
darwin_1872_01
as not
darwin_1859_01 darwin_1860_01 darwin_1872_01
to have
darwin_1859_01 darwin_1860_01
manufactured pottery existed in the valley of the Nile thirteen or fourteen thousand years ago; and who will pretend to say how long before these ancient periods, savages, like those of Tierra del Fuego or Australia, who possess a semi-
darwin_1859_01 darwin_1860_01 darwin_1872_01
domestic
darwin_1859_01 darwin_1860_01
dog, m
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
y not have exis
darwin_1859_01 darwin_1860_01 darwin_1872_01
ted
darwin_1859_01 darwin_1860_01
in
darwin_1872_01
at
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
E
darwin_1872_01
least the do
darwin_1859_01 darwin_1860_01 darwin_1872_01
g
darwin_1859_01 darwin_1860_01
ypt?
darwin_1872_01
.
darwin_1859_01 darwin_1860_01 darwin_1872_01
The
darwin_1859_01 darwin_1860_01
wh
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
le subject must, I think, remain vague; neverthel
darwin_1859_01
s
darwin_1860_01
e
darwin_1859_01 darwin_1860_01
ss, I may, without here ente
darwin_1859_01 darwin_1860_01 darwin_1872_01
ri
darwin_1859_01 darwin_1860_01
n
darwin_1859_01 darwin_1860_01 darwin_1872_01
g
darwin_1859_01 darwin_1860_01
on any details, state that, from geographical and other considerations, I th
darwin_1859_01 darwin_1860_01 darwin_1872_01
in
darwin_1859_01 darwin_1860_01
k
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
i
darwin_1872_01
of mos
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
highly pr
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
bable that
darwin_1872_01
f
darwin_1859_01 darwin_1860_01 darwin_1872_01
our domestic
darwin_1872_01
animals will probably for ever remain vague. But I may here state, that, looking to the domestic
darwin_1859_01 darwin_1860_01 darwin_1872_01
dogs
darwin_1872_01
of the whole world, I
darwin_1859_01 darwin_1860_01 darwin_1872_01
have
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
d
darwin_1872_01
aft
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
r a laboriou
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
c
darwin_1872_01
oll
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
ctio
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1859_01 darwin_1860_01
ded
darwin_1872_01
of all known
darwin_1859_01 darwin_1860_01 darwin_1872_01
f
darwin_1859_01 darwin_1860_01
r
darwin_1872_01
acts, c
darwin_1859_01 darwin_1860_01 darwin_1872_01
om
darwin_1872_01
e to the conclusion that
darwin_1859_01 darwin_1860_01 darwin_1872_01
several wild species
darwin_1860_01
. Knowing, as we do, that savages are very fond
darwin_1860_01 darwin_1872_01
of
darwin_1860_01
taming
darwin_1872_01
C
darwin_1860_01 darwin_1872_01
ani
darwin_1860_01
mals, it seems to me unlikely, in the case of the
darwin_1860_01 darwin_1872_01
d
darwin_1860_01
og-genus, which is distributed in a wild state throughout the world, that since man first appeared one single species alone should
darwin_1872_01
æ
darwin_1860_01 darwin_1872_01
have been
darwin_1872_01
tamed, and that their blood, in some cases mingled together, flows in the veins of our
darwin_1860_01 darwin_1872_01
domestic
darwin_1860_01
at
darwin_1872_01
bre
darwin_1860_01 darwin_1872_01
ed
darwin_1872_01
s
darwin_1860_01 darwin_1872_01 darwin_1859_01
. In regard to sheep and goats I can form no
darwin_1872_01
decided
darwin_1859_01 darwin_1860_01 darwin_1872_01
opinion.
darwin_1859_01 darwin_1860_01
I should think, f
darwin_1872_01
F
darwin_1859_01 darwin_1860_01 darwin_1872_01
rom facts communicated to me by Mr. Blyth, on the habits, voice,
darwin_1859_01 darwin_1860_01
and
darwin_1859_01 darwin_1860_01 darwin_1872_01
constitution,
darwin_1859_01 darwin_1860_01
&
darwin_1872_01
and stru
darwin_1859_01 darwin_1860_01 darwin_1872_01
c
darwin_1859_01 darwin_1860_01
.,
darwin_1872_01
ture
darwin_1859_01 darwin_1860_01 darwin_1872_01
of the humped Indian cattle,
darwin_1872_01
it is almost certain
darwin_1859_01 darwin_1860_01 darwin_1872_01
that the
darwin_1859_01 darwin_1860_01
se
darwin_1872_01
y
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
h
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
d
darwin_1872_01
re
darwin_1859_01 darwin_1860_01 darwin_1872_01
descended from a different aboriginal stock from our European cattle; and s
darwin_1872_01
om
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
veral
darwin_1859_01 darwin_1860_01 darwin_1872_01
competent judges believe that these latter have had
darwin_1859_01 darwin_1860_01
m
darwin_1872_01
two
darwin_1859_01 darwin_1860_01 darwin_1872_01
or
darwin_1859_01 darwin_1860_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
th
darwin_1859_01 darwin_1860_01
an on
darwin_1872_01
re
darwin_1859_01 darwin_1860_01 darwin_1872_01
e wild p
darwin_1872_01
rogenitors,—whether or not these deserve to be called species. This conclusion, as well as that of the specific distinction between the humped and common cattle, may, indeed, be looked upon as established by the admirable rese
darwin_1859_01 darwin_1860_01 darwin_1872_01
ar
darwin_1872_01
ch
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
n
darwin_1872_01
s of Professor Rü
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1872_01
imeyer
darwin_1859_01 darwin_1860_01 darwin_1872_01
. With respect to horses, from reasons which I cannot
darwin_1859_01 darwin_1860_01
give
darwin_1859_01 darwin_1860_01 darwin_1872_01
her
darwin_1872_01
e giv
darwin_1859_01 darwin_1860_01 darwin_1872_01
e, I am doubtfully inclined to believe, in opposition to several authors, that all the races
darwin_1859_01 darwin_1860_01
hav
darwin_1872_01
b
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
descended from
darwin_1872_01
l
darwin_1859_01 darwin_1860_01 darwin_1872_01
on
darwin_1859_01 darwin_1860_01
e
darwin_1872_01
g
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
wild s
darwin_1859_01 darwin_1860_01 darwin_1872_01
to
darwin_1859_01 darwin_1860_01
ck. Mr. Blyth, whose opinion, from his large and varied stores of knowledge, I should value more
darwin_1859_01 darwin_1860_01 darwin_1872_01
th
darwin_1859_01 darwin_1860_01
an that of almost any on
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
th
darwin_1872_01
same species. Hav
darwin_1859_01 darwin_1860_01 darwin_1872_01
in
darwin_1872_01
g
darwin_1859_01 darwin_1860_01 darwin_1872_01
k
darwin_1859_01 darwin_1860_01
s
darwin_1872_01
ept
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
th
darwin_1872_01
ne
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
t
darwin_1872_01
rly
darwin_1859_01 darwin_1860_01 darwin_1872_01
all the
darwin_1872_01
English
darwin_1859_01 darwin_1860_01 darwin_1872_01
breeds of
darwin_1859_01 darwin_1860_01
p
darwin_1872_01
the f
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
u
darwin_1872_01
w
darwin_1859_01 darwin_1860_01 darwin_1872_01
l
darwin_1859_01 darwin_1860_01
try
darwin_1872_01
alive,
darwin_1859_01 darwin_1860_01 darwin_1872_01
hav
darwin_1872_01
ing bred and crossed them, and examined their skeletons, it appears to m
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
pr
darwin_1872_01
alm
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1872_01
st
darwin_1859_01 darwin_1860_01 darwin_1872_01
ce
darwin_1872_01
rtain that all ar
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
ded from
darwin_1859_01 darwin_1860_01 darwin_1872_01
the
darwin_1872_01
des
darwin_1859_01 darwin_1860_01 darwin_1872_01
c
darwin_1872_01
endants
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
mmon
darwin_1872_01
f the
darwin_1859_01 darwin_1860_01 darwin_1872_01
wild Indian fowl
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
(
darwin_1859_01 darwin_1860_01 darwin_1872_01
Gallus bankiva
darwin_1859_01 darwin_1860_01
)
darwin_1872_01
; and this is the conclusion of Mr. Blyth, and of others who have studied this bird in India
darwin_1859_01 darwin_1860_01 darwin_1872_01
. In regard to ducks and rabbits,
darwin_1859_01 darwin_1860_01
th
darwin_1872_01
som
darwin_1859_01 darwin_1860_01 darwin_1872_01
e breeds of which differ
darwin_1872_01
mu
darwin_1859_01 darwin_1860_01 darwin_1872_01
c
darwin_1859_01 darwin_1860_01
onsiderably
darwin_1872_01
h
darwin_1859_01 darwin_1860_01 darwin_1872_01
from each other
darwin_1859_01 darwin_1860_01
in structure
darwin_1859_01 darwin_1860_01 darwin_1872_01
,
darwin_1859_01 darwin_1860_01
I do no
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1872_01
he
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1872_01
evi
darwin_1859_01 darwin_1860_01 darwin_1872_01
d
darwin_1859_01 darwin_1860_01
oubt
darwin_1872_01
ence is clear
darwin_1859_01 darwin_1860_01 darwin_1872_01
that they
darwin_1859_01
all
darwin_1859_01 darwin_1860_01
h
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
v
darwin_1872_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1860_01 darwin_1872_01
all
darwin_1859_01 darwin_1860_01 darwin_1872_01
descended from the common wild duck and rabbit. The doctrine of the origin of our several domestic races from several aboriginal stocks, has been carried to an absurd extreme by some authors. They believe that every race which breeds true, let the distinctive characters be ever so slight, has had its wild prototype. At this rate there must have existed at least a score of species of wild cattle, as many sheep, and several goats
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
in Europe alone, and several even within Great Britain. One author believes that there formerly existed
darwin_1859_01 darwin_1860_01
in Great Britain
darwin_1859_01 darwin_1860_01 darwin_1872_01
eleven wild species of sheep peculiar to
darwin_1872_01
Great Br
darwin_1859_01 darwin_1860_01 darwin_1872_01
it
darwin_1872_01
ain
darwin_1859_01 darwin_1860_01 darwin_1872_01
! When we bear in mind that Britain has now
darwin_1859_01 darwin_1860_01
hardly
darwin_1872_01
not
darwin_1859_01 darwin_1860_01 darwin_1872_01
one peculiar mammal, and France but few distinct from those of Germany
darwin_1859_01 darwin_1860_01
and conversely
darwin_1859_01 darwin_1860_01 darwin_1872_01
, and so with Hungary, Spain, &c., but that each of these kingdoms possesses several peculiar breeds of cattle, sheep, c., we must admit that many domestic breeds
darwin_1872_01
must
darwin_1859_01 darwin_1860_01 darwin_1872_01
have originated in Europe; for whence
darwin_1872_01
otherwise
darwin_1859_01 darwin_1860_01 darwin_1872_01
could they have been derived
darwin_1859_01 darwin_1860_01
, as these several countries do not possess a number of peculiar species as distinct parent-stocks
darwin_1859_01 darwin_1860_01 darwin_1872_01
? So it is in India. Even in the case of the
darwin_1872_01
breeds of the
darwin_1859_01 darwin_1860_01 darwin_1872_01
domestic dog
darwin_1859_01 darwin_1860_01
s of
darwin_1859_01 darwin_1860_01 darwin_1872_01
th
darwin_1859_01 darwin_1860_01
e w
darwin_1872_01
roug
darwin_1859_01 darwin_1860_01 darwin_1872_01
ho
darwin_1859_01 darwin_1860_01
l
darwin_1872_01
ut th
darwin_1859_01 darwin_1860_01 darwin_1872_01
e world, which I
darwin_1859_01 darwin_1860_01
fully
darwin_1859_01 darwin_1860_01 darwin_1872_01
admit
darwin_1859_01 darwin_1860_01
h
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
ve p
darwin_1859_01 darwin_1860_01 darwin_1872_01
r
darwin_1859_01 darwin_1860_01
obably
darwin_1872_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
descended from several wild species,
darwin_1859_01 darwin_1860_01
I
darwin_1872_01
it
darwin_1859_01 darwin_1860_01 darwin_1872_01
cannot
darwin_1872_01
be
darwin_1859_01 darwin_1860_01 darwin_1872_01
doubt
darwin_1872_01
ed
darwin_1859_01 darwin_1860_01 darwin_1872_01
that there has been an immense amount of inherited variation
darwin_1859_01 darwin_1860_01
.
darwin_1872_01
;
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
W
darwin_1872_01
for w
darwin_1859_01 darwin_1860_01 darwin_1872_01
ho
darwin_1859_01 darwin_1860_01
can
darwin_1872_01
will
darwin_1859_01 darwin_1860_01 darwin_1872_01
believe that animals closely resembling the Italian greyhound, the bloodhound, the bull-dog,
darwin_1872_01
pug-dog,
darwin_1859_01 darwin_1860_01 darwin_1872_01
or Blenheim spaniel, &c.—so unlike all wild Canidæ—ever existed
darwin_1859_01 darwin_1860_01
freely
darwin_1859_01 darwin_1860_01 darwin_1872_01
in a state of nature? It has often been loosely said that all our races of dogs have been produced by the crossing of a few aboriginal species; but by crossing we can
darwin_1859_01
get
darwin_1859_01 darwin_1860_01 darwin_1872_01
only
darwin_1860_01 darwin_1872_01
get
darwin_1859_01 darwin_1860_01 darwin_1872_01
forms in some degree intermediate between their parents; and if we account for our several domestic races by this process, we must admit the former existence of the most extreme forms, as the Italian greyhound, bloodhound, bull-dog, &c., in the wild state. Moreover, the possibility of making distinct races by crossing has been greatly exaggerated.
darwin_1859_01 darwin_1860_01
Th
darwin_1872_01
Many cas
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
s a
darwin_1859_01 darwin_1860_01 darwin_1872_01
re
darwin_1859_01 darwin_1860_01
ca
darwin_1872_01
o
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1859_01 darwin_1860_01
b
darwin_1872_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
cord,
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
n
darwin_1872_01
sh
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
doubt
darwin_1872_01
wing
darwin_1859_01 darwin_1860_01 darwin_1872_01
that a race may be modified by occasional crosses, if aided by the careful selection of th
darwin_1859_01 darwin_1860_01
os
darwin_1859_01 darwin_1860_01 darwin_1872_01
e individual
darwin_1859_01 darwin_1860_01
mongrel
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
which present
darwin_1859_01 darwin_1860_01
any
darwin_1872_01
the
darwin_1859_01 darwin_1860_01 darwin_1872_01
desired character; but t
darwin_1859_01 darwin_1860_01
hat a race c
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
uld be
darwin_1859_01 darwin_1860_01 darwin_1872_01
obtain
darwin_1859_01 darwin_1860_01
ed
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
n
darwin_1872_01
a rac
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
arly
darwin_1859_01 darwin_1860_01 darwin_1872_01
intermediate between two
darwin_1859_01 darwin_1860_01
ex
darwin_1872_01
qui
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
mely
darwin_1859_01 darwin_1860_01 darwin_1872_01
di
darwin_1859_01 darwin_1860_01
ffere
darwin_1872_01
sti
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1872_01
c
darwin_1859_01 darwin_1860_01 darwin_1872_01
t races
darwin_1859_01 darwin_1860_01
or spec
darwin_1859_01
e
darwin_1859_01 darwin_1860_01
ies
darwin_1859_01 darwin_1860_01 darwin_1872_01
,
darwin_1859_01 darwin_1860_01
I can hard
darwin_1872_01
wou
darwin_1859_01 darwin_1860_01 darwin_1872_01
l
darwin_1859_01 darwin_1860_01
y
darwin_1872_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
be
darwin_1859_01 darwin_1860_01
lie
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
ve
darwin_1872_01
ry difficult
darwin_1859_01 darwin_1860_01 darwin_1872_01
. Sir J. Sebright expressly experiment
darwin_1859_01 darwin_1860_01
is
darwin_1859_01 darwin_1860_01 darwin_1872_01
ed
darwin_1859_01 darwin_1860_01
for
darwin_1872_01
with
darwin_1859_01 darwin_1860_01 darwin_1872_01
this object, and failed. The offspring from the first cross between two pure breeds is tolerably and sometimes (as I have found with pigeons)
darwin_1859_01 darwin_1860_01
ex
darwin_1872_01
qui
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
mely
darwin_1859_01 darwin_1860_01 darwin_1872_01
uniform
darwin_1872_01
in character
darwin_1859_01 darwin_1860_01 darwin_1872_01
, and everything seems simple enough; but when these mongrels are crossed one with another for several generations, hardly two of them
darwin_1859_01 darwin_1860_01
will b
darwin_1872_01
ar
darwin_1859_01 darwin_1860_01 darwin_1872_01
e alike, and then the
darwin_1859_01 darwin_1860_01
extreme
darwin_1859_01 darwin_1860_01 darwin_1872_01
difficulty
darwin_1859_01 darwin_1860_01
, or rather utter hopelessness,
darwin_1859_01 darwin_1860_01 darwin_1872_01
of the task becomes
darwin_1859_01 darwin_1860_01
apparent. Certainly, a breed intermediate between two very distinct breeds could not be got without extreme care and long-continued selection; nor can I find a single case on record of a per
darwin_1859_01 darwin_1860_01 darwin_1872_01
man
darwin_1859_01 darwin_1860_01
ent race hav
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
ng been thus
darwin_1859_01 darwin_1860_01 darwin_1872_01
f
darwin_1859_01 darwin_1860_01
orm
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
d. On
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
he
darwin_1872_01
.
darwin_1859_01 darwin_1860_01 darwin_1872_01
Breeds of the Domestic Pigeon
darwin_1872_01
, their Differences and Origin
darwin_1859_01 darwin_1860_01 darwin_1872_01
.
darwin_1859_01 darwin_1860_01
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
Believing that it is always best to study some special group, I have, after deliberation, taken up domestic pigeons. I have kept every breed which I could purchase or obtain, and have been most kindly favoured with skins from several quarters of the world, more especially by the Hon. W. Elliot
darwin_1872_01
t
darwin_1859_01 darwin_1860_01 darwin_1872_01
from India, and by the Hon. C. Murray from Persia. Many treatises in different languages have been published on pigeons, and some of them are very important, as being of considerable antiquity. I have associated with several eminent fanciers, and have been permitted to join two of the London Pigeon Clubs. The diversity of the breeds is something astonishing. Compare the English carrier and the short-faced tumbler, and see the wonderful difference in their beaks, entailing corresponding differences in their skulls. The carrier, more especially the male bird, is also remarkable from the wonderful development of the carunculated skin about the head
darwin_1859_01 darwin_1860_01
,
darwin_1872_01
;
darwin_1859_01 darwin_1860_01 darwin_1872_01
and this is accompanied by greatly elongated eyelids, very large external orifices to the nostrils, and a wide gape of mouth. The short-faced tumbler has a beak in outline almost like that of a finch; and the common tumbler has the singular
darwin_1859_01
and strictly
darwin_1859_01 darwin_1860_01 darwin_1872_01
inherited habit of flying at a great height in a compact flock, and tumbling in the air head over heels. The runt is a bird of great size, with long
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
massive beak and large feet; some of the sub-breeds of runts have very long necks, others very long wings and tails, others singularly short tails. The barb is allied to the carrier, but, instead of a
darwin_1859_01 darwin_1860_01
very
darwin_1859_01 darwin_1860_01 darwin_1872_01
long beak, has a very short and
darwin_1859_01 darwin_1860_01
very
darwin_1859_01 darwin_1860_01 darwin_1872_01
broad one. The pouter has a much elongated body, wings, and legs; and its enormously developed crop, which it glories in inflating, may well excite astonishment and even laughter. The turbit has a
darwin_1859_01 darwin_1860_01
very
darwin_1859_01 darwin_1860_01 darwin_1872_01
short and conical beak, with a line of reversed feathers down the breast; and it has the habit of continually expanding
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
slightly
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
the upper part of the œsophagus. The Jacobin has the feathers so much reversed along the back of the neck that they form a hood
darwin_1859_01 darwin_1860_01
,
darwin_1872_01
;
darwin_1859_01 darwin_1860_01 darwin_1872_01
and it has, proportionally to its size,
darwin_1859_01 darwin_1860_01
much
darwin_1859_01 darwin_1860_01 darwin_1872_01
elongated wing and tail feathers. The trumpeter and laugher, as their names express, utter a very different coo from the other breeds. The fantail has thirty or even forty tail
darwin_1859_01 darwin_1872_01
-
darwin_1860_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
feathers, instead of twelve or fourteen
darwin_1859_01 darwin_1860_01
,
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
the normal number in all
darwin_1872_01
the
darwin_1859_01 darwin_1860_01 darwin_1872_01
members of the great pigeon family
darwin_1859_01 darwin_1860_01
; and
darwin_1872_01
:
darwin_1859_01 darwin_1860_01 darwin_1872_01
these feathers are kept expanded, and are carried so erect
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
that in good birds the head and tail touch
darwin_1859_01 darwin_1860_01
;
darwin_1872_01
:
darwin_1859_01 darwin_1860_01 darwin_1872_01
the oil-gland is quite aborted. Several other less distinct breeds might
darwin_1859_01
have
darwin_1859_01 darwin_1860_01 darwin_1872_01
be
darwin_1859_01
en
darwin_1859_01 darwin_1860_01 darwin_1872_01
specified. In the skeletons of the several breeds, the development of the bones of the face in length and breadth and curvature differs enormously. The shape, as well as the breadth and length of the ramus of the lower jaw, varies in a highly remarkable manner. The
darwin_1859_01 darwin_1860_01
number of the
darwin_1859_01 darwin_1860_01 darwin_1872_01
caudal and sacral vertebræ vary
darwin_1872_01
in number
darwin_1859_01 darwin_1860_01 darwin_1872_01
; as does the number of the ribs, together with their relative breadth and the presence of processes. The size and shape of the apertures in the sternum are highly variable; so is the degree of divergence and relative size of the two arms of the furcula. The proportional width of the gape of mouth, the proportional length of the eyelids, of the orifice of the nostrils, of the tongue (not always in strict correlation with the length of beak), the size of the crop and of the upper part of the œsophagus; the development and abortion of the oil-gland; the number of the primary wing and caudal feathers; the relative length of
darwin_1872_01
the
darwin_1859_01 darwin_1860_01 darwin_1872_01
wing and tail to each other and to the body; the relative length of
darwin_1872_01
the
darwin_1859_01 darwin_1860_01 darwin_1872_01
leg and
darwin_1859_01 darwin_1860_01
o
darwin_1859_01 darwin_1860_01 darwin_1872_01
f
darwin_1859_01 darwin_1860_01
the fee
darwin_1872_01
oo
darwin_1859_01 darwin_1860_01 darwin_1872_01
t; the number of scutellæ on the toes, the development of skin between the toes, are all points of structure which are variable. The period at which the perfect plumage is acquired varies, as does the state of the down with which the nestling birds are clothed when hatched. The shape and size of the eggs vary. The manner of flight
darwin_1859_01 darwin_1860_01
differs remarkably;
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
s
darwin_1872_01
n
darwin_1859_01 darwin_1860_01 darwin_1872_01
d
darwin_1859_01 darwin_1860_01
oes
darwin_1859_01 darwin_1860_01 darwin_1872_01
in some breeds the voice and disposition
darwin_1872_01
, differ remarkably
darwin_1859_01 darwin_1860_01 darwin_1872_01
. Lastly, in certain breeds, the males and females have come to differ
darwin_1859_01 darwin_1860_01
to
darwin_1872_01
in
darwin_1859_01 darwin_1860_01 darwin_1872_01
a slight degree from each other. Altogether at least a score of pigeons might be chosen, which
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
if shown to an ornithologist, and he were told that they were wild birds, would certainly
darwin_1859_01 darwin_1860_01
, I think,
darwin_1859_01 darwin_1860_01 darwin_1872_01
be ranked by him as well-defined species. Moreover, I do not believe that any ornithologist would
darwin_1872_01
in this case
darwin_1859_01 darwin_1860_01 darwin_1872_01
place the English carrier, the short-faced tumbler, the runt, the barb, pouter, and fantail in the same genus; more especially as in each of these breeds several truly-inherited sub-breeds, or species
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
as he
darwin_1859_01 darwin_1860_01
might have
darwin_1872_01
would
darwin_1859_01 darwin_1860_01 darwin_1872_01
call
darwin_1859_01 darwin_1860_01
ed
darwin_1859_01 darwin_1860_01 darwin_1872_01
them, could be shown him. Great as
darwin_1872_01
are
darwin_1859_01 darwin_1860_01 darwin_1872_01
the differences
darwin_1859_01 darwin_1860_01
are
darwin_1859_01 darwin_1860_01 darwin_1872_01
between the breeds of
darwin_1872_01
the
darwin_1859_01 darwin_1860_01 darwin_1872_01
pigeon
darwin_1859_01 darwin_1860_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
, I am fully convinced that the common opinion of naturalists is correct, namely, that all
darwin_1859_01 darwin_1860_01
h
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
v
darwin_1872_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
e descended from the rock
darwin_1859_01 darwin_1860_01
-
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
pigeon (Columba livia), including under this term several geographical races or sub-species, which differ from each other in the most trifling respects. As several of the reasons which have led me to this belief are in some degree applicable in other cases, I will here briefly give them. If the several breeds are not varieties, and have not proceeded from the rock-pigeon, they must have descended from at least seven or eight aboriginal stocks; for it is impossible to make the present domestic breeds by the crossing of any lesser number: how, for instance, could a pouter be produced by crossing two breeds unless one of the parent-stocks possessed the characteristic enormous crop? The supposed aboriginal stocks must all have been rock-pigeons, that is,
darwin_1872_01
they did
darwin_1859_01 darwin_1860_01 darwin_1872_01
not breed
darwin_1859_01 darwin_1860_01
ing
darwin_1859_01 darwin_1860_01 darwin_1872_01
or willingly perch
darwin_1859_01 darwin_1860_01
ing
darwin_1859_01 darwin_1860_01 darwin_1872_01
on trees. But besides C. livia, with its geographical sub-species, only two or three other species of rock-pigeons are known; and these have not any of the characters of the domestic breeds. Hence the supposed aboriginal stocks must either still exist in the countries where they were originally domesticated, and yet be unknown to ornithologists; and this, considering their size, habits, and remarkable characters, seems
darwin_1859_01 darwin_1860_01
very
darwin_1859_01 darwin_1860_01 darwin_1872_01
improbable; or they must have become extinct in the wild state. But birds breeding on precipices, and good fliers, are unlikely to be exterminated; and the common rock-pigeon, which has the same habits with the domestic breeds, has not been exterminated even on several of the smaller British islets, or on the shores of the Mediterranean. Hence the supposed extermination of so many species having similar habits with the rock-pigeon seems
darwin_1859_01 darwin_1860_01
to me
darwin_1859_01 darwin_1860_01 darwin_1872_01
a very rash assumption. Moreover, the several above-named domesticated breeds have been transported to all parts of the world, and, therefore, some of them must have been carried back again into their native country; but not one has
darwin_1859_01 darwin_1860_01
ever
darwin_1859_01 darwin_1860_01 darwin_1872_01
become wild or feral, though the dovecot-pigeon, which is the rock-pigeon in a very slightly altered state, has become feral in several places. Again, all recent experience shows that it is
darwin_1859_01 darwin_1860_01
most
darwin_1859_01 darwin_1860_01 darwin_1872_01
difficult to get
darwin_1859_01 darwin_1860_01
any
darwin_1859_01 darwin_1860_01 darwin_1872_01
wild animal
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
to breed freely under domestication; yet
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
on the hypothesis of the multiple origin of our pigeons, it must be assumed that at least seven or eight species were so thoroughly domesticated in ancient times by half-civili
darwin_1859_01 darwin_1860_01
z
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
ed man, as to be quite prolific under confinement. An argument
darwin_1859_01 darwin_1860_01
, as it seems to me,
darwin_1859_01 darwin_1860_01 darwin_1872_01
of great weight, and applicable in several other cases, is, that the above-specified breeds, though agreeing generally
darwin_1872_01
with the wild rock-pigeon
darwin_1859_01 darwin_1860_01 darwin_1872_01
in constitution, habits, voice, colouring, and in most parts of their structure,
darwin_1859_01 darwin_1860_01
with the wild rock-pigeon,
darwin_1859_01 darwin_1860_01 darwin_1872_01
yet are certainly highly abnormal in other parts
darwin_1859_01 darwin_1860_01
of their structure
darwin_1859_01
:
darwin_1860_01 darwin_1872_01
;
darwin_1859_01 darwin_1860_01 darwin_1872_01
we may look in vain through
darwin_1859_01 darwin_1860_01
out
darwin_1859_01 darwin_1860_01 darwin_1872_01
the whole great family of Columbidæ for a beak like that of the English carrier, or that of the short-faced tumbler, or barb; for reversed feathers like those of the
darwin_1860_01 darwin_1872_01
J
darwin_1859_01
j
darwin_1859_01 darwin_1860_01 darwin_1872_01
acobin; for a crop like that of the pouter; for tail-feathers like those of the fantail. Hence it must be assumed not only that half-civili
darwin_1859_01 darwin_1860_01
z
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
ed man succeeded in thoroughly domesticating several species, but that he intentionally or by chance picked out extraordinarily abnormal species; and further, that these very species have since all become extinct or unknown. So many strange contingencies
darwin_1859_01 darwin_1860_01
seem to m
darwin_1872_01
ar
darwin_1859_01 darwin_1860_01 darwin_1872_01
e improbable in the highest degree. Some facts in regard to the colouring of pigeons well deserve consideration. The rock
darwin_1859_01 darwin_1860_01
-
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
pigeon is of a slaty-blue,
darwin_1859_01 darwin_1860_01
and
darwin_1872_01
wit
darwin_1859_01 darwin_1860_01 darwin_1872_01
h
darwin_1859_01 darwin_1860_01
as a
darwin_1859_01 darwin_1860_01 darwin_1872_01
white
darwin_1859_01 darwin_1860_01
r
darwin_1872_01
loins; b
darwin_1859_01 darwin_1860_01 darwin_1872_01
u
darwin_1859_01 darwin_1860_01
mp
darwin_1872_01
t
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
(
darwin_1859_01 darwin_1860_01 darwin_1872_01
the Indian sub-species, C. intermedia of Strickland, ha
darwin_1859_01 darwin_1860_01
v
darwin_1872_01
s th
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
ng
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
i
darwin_1872_01
par
darwin_1859_01 darwin_1860_01 darwin_1872_01
t bluish
darwin_1859_01 darwin_1860_01
);
darwin_1872_01
.
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
t
darwin_1872_01
T
darwin_1859_01 darwin_1860_01 darwin_1872_01
he tail has a terminal dark bar, with the
darwin_1859_01 darwin_1860_01
bases of the
darwin_1859_01 darwin_1860_01 darwin_1872_01
outer feathers externally edged
darwin_1872_01
at the base
darwin_1859_01 darwin_1860_01 darwin_1872_01
with white
darwin_1859_01 darwin_1860_01
;
darwin_1872_01
.
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
t
darwin_1872_01
T
darwin_1859_01 darwin_1860_01 darwin_1872_01
he wings have two black bars
darwin_1859_01 darwin_1860_01
;
darwin_1872_01
.
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
s
darwin_1872_01
S
darwin_1859_01 darwin_1860_01 darwin_1872_01
ome semi-domestic breeds
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
and some
darwin_1859_01 darwin_1860_01
apparently
darwin_1859_01 darwin_1860_01 darwin_1872_01
truly wild breeds
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
have, besides the two black bars, the wings chequered with black. These several marks do not occur together in any other species of the whole family. Now, in every one of the domestic breeds, taking thoroughly well-bred birds, all the above marks, even to the white edging of the outer tail-feathers, sometimes concur perfectly developed. Moreover, when
darwin_1859_01 darwin_1860_01
two
darwin_1859_01 darwin_1860_01 darwin_1872_01
birds belonging to two
darwin_1872_01
or more
darwin_1859_01 darwin_1860_01 darwin_1872_01
distinct breeds are crossed, n
darwin_1872_01
on
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
ither
darwin_1859_01 darwin_1860_01 darwin_1872_01
of which
darwin_1859_01 darwin_1860_01
is
darwin_1872_01
are
darwin_1859_01 darwin_1860_01 darwin_1872_01
blue or ha
darwin_1859_01 darwin_1860_01
s
darwin_1872_01
ve
darwin_1859_01 darwin_1860_01 darwin_1872_01
any of the above-specified marks, the mongrel offspring are very apt suddenly to acquire these characters
darwin_1859_01 darwin_1860_01
;
darwin_1872_01
.
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
f
darwin_1872_01
T
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
r
darwin_1872_01
give one
darwin_1859_01 darwin_1860_01 darwin_1872_01
instance
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1872_01
out of several which I have observed:—
darwin_1859_01 darwin_1860_01 darwin_1872_01
I crossed some
darwin_1859_01 darwin_1860_01
uniformly
darwin_1859_01 darwin_1860_01 darwin_1872_01
white fantails
darwin_1872_01
, which breed very true,
darwin_1859_01 darwin_1860_01 darwin_1872_01
with some
darwin_1859_01 darwin_1860_01
uniformly
darwin_1859_01 darwin_1860_01 darwin_1872_01
black barbs
darwin_1872_01
—and it so happens that blue varieties of barbs are so rare that I never heard of an instance in England; and the mongrels were black, brown, and mottled. I also crossed a barb with a spot, which is a white bird with a red tail and red spot on the forehead, and which notoriously breeds very true; the mongrels were dusky and mottled. I then crossed one of the mongrel barb-fantails with a mongrel barb-spot
darwin_1859_01 darwin_1860_01 darwin_1872_01
, and they produced
darwin_1859_01 darwin_1860_01
mottled brown
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
nd black
darwin_1859_01 darwin_1860_01 darwin_1872_01
bird
darwin_1859_01 darwin_1860_01
s; these I again crossed together, and one grandchild of the pure white fantail and pure black barb was
darwin_1859_01 darwin_1860_01 darwin_1872_01
of as beautiful a blue colour, with the white
darwin_1859_01 darwin_1860_01
rump
darwin_1872_01
loins
darwin_1859_01 darwin_1860_01 darwin_1872_01
, double black wing-bar, and barred and white-edged tail-feathers, as any wild rock
darwin_1859_01 darwin_1860_01
-
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
pigeon! We can understand these facts, on the well-known principle of reversion to ancestral characters, if all the domestic breeds
darwin_1859_01 darwin_1860_01
h
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
v
darwin_1872_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
e descended from the rock
darwin_1859_01 darwin_1860_01
-
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
pigeon. But if we deny this, we must make one of the two following highly improbable suppositions. Either, first
darwin_1859_01 darwin_1860_01
ly
darwin_1859_01 darwin_1860_01 darwin_1872_01
, that all the several imagined aboriginal stocks were coloured and marked like the rock
darwin_1859_01 darwin_1860_01
-
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
pigeon, although no other existing species is thus coloured and marked, so that in each separate breed there might be a tendency to revert to the very same colours and markings. Or, secondly, that each breed, even the purest, has within a dozen
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
or
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
at most
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
within a score
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
of generations, been crossed by the rock
darwin_1859_01 darwin_1860_01
-
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
pigeon
darwin_1859_01 darwin_1860_01
:
darwin_1872_01
;
darwin_1859_01 darwin_1860_01 darwin_1872_01
I say within a dozen or twenty generations, for
darwin_1859_01 darwin_1860_01
w
darwin_1872_01
no instanc
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
is
darwin_1859_01 darwin_1860_01 darwin_1872_01
know
darwin_1872_01
n
darwin_1859_01 darwin_1860_01 darwin_1872_01
of
darwin_1859_01 darwin_1860_01
no fa
darwin_1859_01 darwin_1860_01 darwin_1872_01
c
darwin_1859_01 darwin_1860_01
t c
darwin_1872_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
unt
darwin_1872_01
ss
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
nancing the belief that the chil
darwin_1859_01 darwin_1860_01 darwin_1872_01
d
darwin_1872_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
v
darwin_1872_01
sc
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
r
darwin_1872_01
ndants
darwin_1859_01 darwin_1860_01 darwin_1872_01
revert
darwin_1859_01 darwin_1860_01
s
darwin_1872_01
ing
darwin_1859_01 darwin_1860_01 darwin_1872_01
to
darwin_1859_01 darwin_1860_01
some o
darwin_1872_01
a
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1859_01 darwin_1860_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
ancestor
darwin_1872_01
of foreign blood
darwin_1859_01 darwin_1860_01 darwin_1872_01
, removed by a greater number of generations. In a breed which has been crossed only once
darwin_1859_01 darwin_1860_01
with some distinct breed
darwin_1859_01 darwin_1860_01 darwin_1872_01
, the tendency to rever
darwin_1859_01 darwin_1860_01
sion
darwin_1872_01
t
darwin_1859_01 darwin_1860_01 darwin_1872_01
to any character derived from such
darwin_1872_01
a
darwin_1859_01 darwin_1860_01 darwin_1872_01
cross will naturally become less and less, as in each succeeding generation there will be less of the foreign blood; but when there has been no cross
darwin_1859_01 darwin_1860_01
with a distinct breed
darwin_1859_01 darwin_1860_01 darwin_1872_01
, and there is a tendency in
darwin_1859_01 darwin_1860_01
bo
darwin_1859_01 darwin_1860_01 darwin_1872_01
th
darwin_1872_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
pa
darwin_1872_01
b
darwin_1859_01 darwin_1860_01 darwin_1872_01
re
darwin_1859_01 darwin_1860_01
nts
darwin_1872_01
ed
darwin_1859_01 darwin_1860_01 darwin_1872_01
to revert to a character
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
which
darwin_1859_01 darwin_1860_01
h
darwin_1872_01
w
darwin_1859_01 darwin_1860_01 darwin_1872_01
as
darwin_1859_01 darwin_1860_01
been
darwin_1859_01 darwin_1860_01 darwin_1872_01
lost during some former generation, this tendency, for all that we can see to the contrary, may be transmitted undiminished for an indefinite number of generations. These two distinct cases
darwin_1872_01
of reversion
darwin_1859_01 darwin_1860_01 darwin_1872_01
are often confounded
darwin_1859_01 darwin_1860_01
in
darwin_1872_01
together by
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
reati
darwin_1872_01
ho
darwin_1859_01 darwin_1860_01 darwin_1872_01
se
darwin_1859_01 darwin_1860_01
s
darwin_1872_01
who have written
darwin_1859_01 darwin_1860_01 darwin_1872_01
on inheritance. Lastly, the hybrids or mongrels from between all the
darwin_1859_01 darwin_1860_01
domestic
darwin_1859_01 darwin_1860_01 darwin_1872_01
breeds of
darwin_1872_01
the
darwin_1859_01 darwin_1860_01 darwin_1872_01
pigeon
darwin_1859_01 darwin_1860_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
are perfectly fertile
darwin_1859_01 darwin_1860_01
.
darwin_1872_01
, as
darwin_1859_01 darwin_1860_01 darwin_1872_01
I can state
darwin_1859_01 darwin_1860_01
this
darwin_1859_01 darwin_1860_01 darwin_1872_01
from my own observations, purposely made
darwin_1860_01 darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
on the most distinct breeds. Now,
darwin_1859_01 darwin_1860_01
it is difficult, per
darwin_1859_01 darwin_1860_01 darwin_1872_01
h
darwin_1859_01 darwin_1860_01
aps impossible, to bring forw
darwin_1859_01 darwin_1860_01 darwin_1872_01
ard
darwin_1872_01
ly
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
o
darwin_1872_01
a
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1859_01 darwin_1860_01
e
darwin_1872_01
y
darwin_1859_01 darwin_1860_01 darwin_1872_01
case
darwin_1872_01
s have been ascertained with certainty
darwin_1859_01 darwin_1860_01 darwin_1872_01
of
darwin_1859_01 darwin_1860_01
the
darwin_1859_01 darwin_1860_01 darwin_1872_01
hybrid
darwin_1859_01 darwin_1860_01
off
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1859_01 darwin_1860_01
pring
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1872_01
fr
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
f
darwin_1872_01
m
darwin_1859_01 darwin_1860_01 darwin_1872_01
two
darwin_1859_01 darwin_1860_01
an
darwin_1872_01
qu
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
mals cl
darwin_1872_01
t
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
arly
darwin_1859_01 darwin_1860_01 darwin_1872_01
distinct
darwin_1872_01
species of animals
darwin_1859_01 darwin_1860_01 darwin_1872_01
being
darwin_1859_01 darwin_1860_01
themselves
darwin_1859_01 darwin_1860_01 darwin_1872_01
perfectly fertile. Some authors believe that long-continued domestication eliminates this strong tendency to sterility
darwin_1859_01 darwin_1860_01
:
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
f
darwin_1872_01
in species. F
darwin_1859_01 darwin_1860_01 darwin_1872_01
rom the history of the dog
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
I
darwin_1872_01
and of some other domestic animals,
darwin_1859_01 darwin_1860_01 darwin_1872_01
thi
darwin_1872_01
s co
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1859_01 darwin_1860_01
k there
darwin_1872_01
clusion
darwin_1859_01 darwin_1860_01 darwin_1872_01
is
darwin_1859_01 darwin_1860_01
some
darwin_1859_01 darwin_1860_01 darwin_1872_01
probab
darwin_1859_01 darwin_1860_01
i
darwin_1859_01 darwin_1860_01 darwin_1872_01
l
darwin_1859_01 darwin_1860_01
it
darwin_1859_01 darwin_1860_01 darwin_1872_01
y
darwin_1872_01
qu
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
n
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
his hypoth
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
sis
darwin_1872_01
correct
darwin_1859_01 darwin_1860_01 darwin_1872_01
, if applied to species closely related to
darwin_1859_01 darwin_1860_01
g
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
ach o
darwin_1859_01 darwin_1860_01 darwin_1872_01
ther
darwin_1859_01 darwin_1860_01
, though it is unsupported by a single experiment
darwin_1859_01 darwin_1860_01 darwin_1872_01
. But to extend
darwin_1859_01 darwin_1860_01
the hypothes
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
s
darwin_1872_01
t
darwin_1859_01 darwin_1860_01 darwin_1872_01
so far as to suppose that species, aboriginally as distinct as carriers, tumblers, pouters, and fantails now are, should yield offspring perfectly fertile
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
inter se,
darwin_1859_01 darwin_1860_01
seems t
darwin_1872_01
w
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1872_01
uld
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
m
darwin_1872_01
b
darwin_1859_01 darwin_1860_01 darwin_1872_01
e rash in the extreme. From these several reasons, namely,
darwin_1859_01 darwin_1860_01
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
the improbability of man having formerly
darwin_1859_01 darwin_1860_01
got
darwin_1872_01
made
darwin_1859_01 darwin_1860_01 darwin_1872_01
seven or eight supposed species of pigeons to breed freely under domestication;
darwin_1859_01 darwin_1860_01
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
these supposed species being quite unknown in a wild state, and their
darwin_1872_01
not having
darwin_1859_01 darwin_1860_01 darwin_1872_01
becom
darwin_1859_01 darwin_1860_01
ing
darwin_1872_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1872_01
a
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1859_01 darwin_1860_01
o
darwin_1872_01
y
darwin_1859_01 darwin_1860_01 darwin_1872_01
where feral;
darwin_1859_01 darwin_1860_01
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
these species
darwin_1859_01 darwin_1860_01
hav
darwin_1872_01
present
darwin_1859_01 darwin_1860_01 darwin_1872_01
ing
darwin_1872_01
certain
darwin_1859_01 darwin_1860_01 darwin_1872_01
very abnormal character
darwin_1859_01 darwin_1860_01
s in certain respect
darwin_1859_01 darwin_1860_01 darwin_1872_01
s, as compared with all other Columbidæ, though so like
darwin_1859_01 darwin_1860_01
in most other respects to
darwin_1859_01 darwin_1860_01 darwin_1872_01
the rock-pigeon
darwin_1872_01
in most respects
darwin_1859_01 darwin_1860_01 darwin_1872_01
;
darwin_1872_01
—the occasional re-appearance of
darwin_1859_01 darwin_1860_01 darwin_1872_01
the blue colour and various
darwin_1872_01
black
darwin_1859_01 darwin_1860_01 darwin_1872_01
marks
darwin_1859_01 darwin_1860_01
occasionally appearing
darwin_1859_01 darwin_1860_01 darwin_1872_01
in all the breeds, both when kept pure and when crossed;
darwin_1872_01
—and lastly,
darwin_1859_01 darwin_1860_01 darwin_1872_01
the mongrel offspring being perfectly fertile;—from these several reasons, taken together,
darwin_1859_01 darwin_1860_01
I
darwin_1872_01
we may safely
darwin_1859_01 darwin_1860_01 darwin_1872_01
c
darwin_1859_01 darwin_1860_01
an feel n
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
darwin_1872_01
nclu
darwin_1859_01 darwin_1860_01 darwin_1872_01
d
darwin_1859_01 darwin_1860_01
oubt
darwin_1872_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
that all our domestic breeds
darwin_1859_01 darwin_1860_01
h
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
v
darwin_1872_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
e descended from the
darwin_1872_01
rock-pigeon or
darwin_1859_01 darwin_1860_01 darwin_1872_01
Columba livia with its geographical sub-species. In favour of this view, I may add, firstly, that
darwin_1872_01
the wild
darwin_1859_01 darwin_1860_01 darwin_1872_01
C. livia
darwin_1859_01 darwin_1860_01
, or the rock-pigeon,
darwin_1859_01 darwin_1860_01 darwin_1872_01
has been found capable of domestication in Europe and in India; and that it agrees in habits and in a great number of points of structure with all the domestic breeds. Secondly,
darwin_1872_01
that,
darwin_1859_01 darwin_1860_01 darwin_1872_01
although an English carrier or
darwin_1872_01
a
darwin_1859_01 darwin_1860_01 darwin_1872_01
short-faced tumbler differs immensely in certain characters from the rock-pigeon, yet
darwin_1872_01
that,
darwin_1859_01 darwin_1860_01 darwin_1872_01
by comparing the several sub-breeds of these
darwin_1859_01
b
darwin_1860_01
va
darwin_1859_01 darwin_1860_01
r
darwin_1860_01
i
darwin_1859_01 darwin_1860_01
e
darwin_1860_01 darwin_1872_01
t
darwin_1860_01
i
darwin_1872_01
wo rac
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
s, more especially those brought from distant countries, we can make
darwin_1872_01
, between them and the rock pigeon,
darwin_1859_01 darwin_1860_01 darwin_1872_01
an almost perfect series
darwin_1872_01
; so we can in some other cases,
darwin_1859_01 darwin_1860_01 darwin_1872_01
b
darwin_1859_01 darwin_1860_01
e
darwin_1872_01
u
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1872_01
not
darwin_1859_01 darwin_1860_01 darwin_1872_01
w
darwin_1859_01 darwin_1860_01
een
darwin_1872_01
ith all
darwin_1859_01 darwin_1860_01 darwin_1872_01
the
darwin_1859_01 darwin_1860_01
ext
darwin_1872_01
b
darwin_1859_01 darwin_1860_01 darwin_1872_01
re
darwin_1859_01 darwin_1860_01
m
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1859_01 darwin_1860_01
of structure
darwin_1859_01 darwin_1860_01 darwin_1872_01
. Thirdly, those characters which are mainly distinctive of each breed
darwin_1872_01
are in each eminently variable
darwin_1859_01 darwin_1860_01 darwin_1872_01
, for instance the wattle and length of beak of the carrier, the shortness of that of the tumbler, and the number of tail-feathers in the fantail
darwin_1859_01 darwin_1860_01
, are in each breed eminently variable
darwin_1859_01 darwin_1860_01 darwin_1872_01
; and the explanation of this fact will be obvious when we
darwin_1859_01 darwin_1860_01
come to
darwin_1859_01 darwin_1860_01 darwin_1872_01
treat of
darwin_1859_01 darwin_1860_01
s
darwin_1872_01
S
darwin_1859_01 darwin_1860_01 darwin_1872_01
election. Fourthly, pigeons have been watched
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
and tended with the utmost care, and loved by many people. They have been domesticated for thousands of years in several quarters of the world; the earliest known record of pigeons is in the fifth Ægyptian dynasty, about 3000 B.C., as was pointed out to me by Professor Lepsius; but Mr. Birch informs me that pigeons are given in a bill of fare in the previous dynasty. In the time of the Romans, as we hear from Pliny, immense prices were given for pigeons; "nay, they are come to this pass, that they can reckon up their pedigree and race." Pigeons were much valued by Akber Khan in India, about the year 1600; never less than 20,000 pigeons were taken with the court. "The monarchs of Iran and Turan sent him some very rare birds
darwin_1859_01 darwin_1860_01
;
darwin_1859_01 darwin_1860_01 darwin_1872_01
"
darwin_1872_01
;
darwin_1859_01 darwin_1860_01 darwin_1872_01
and, continues the courtly historian, "His Majesty by crossing the breeds, which method was never practised before, has improved them astonishingly." About this same period the Dutch were as eager about pigeons as were the old Romans. The paramount importance of these considerations in explaining the immense amount of variation which pigeons have undergone, will
darwin_1872_01
likewise
darwin_1859_01 darwin_1860_01 darwin_1872_01
be obvious when we treat of Selection. We shall then, also, see how it is that the
darwin_1872_01
several
darwin_1859_01 darwin_1860_01 darwin_1872_01
breeds so often have a somewhat monstrous character. It is also a most favourable circumstance for the production of distinct breeds, that male and female pigeons can be easily mated for life; and thus different breeds can be kept together in the same aviary. I have discussed the probable origin of domestic pigeons at some, yet quite insufficient, length; because when I first kept pigeons and watched the several kinds,
darwin_1872_01
well
darwin_1859_01 darwin_1860_01 darwin_1872_01
knowing
darwin_1859_01 darwin_1860_01
well
darwin_1859_01 darwin_1860_01 darwin_1872_01
how tru
darwin_1859_01 darwin_1860_01
e
darwin_1872_01
ly
darwin_1859_01 darwin_1860_01 darwin_1872_01
they br
darwin_1872_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
ed, I felt fully as much difficulty in believing that
darwin_1872_01
since
darwin_1859_01 darwin_1860_01 darwin_1872_01
they
darwin_1859_01 darwin_1860_01
coul
darwin_1872_01
ha
darwin_1859_01 darwin_1860_01 darwin_1872_01
d
darwin_1859_01
ever
darwin_1872_01
been domesticated they
darwin_1859_01 darwin_1860_01 darwin_1872_01
ha
darwin_1859_01 darwin_1860_01
ve
darwin_1872_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
des
darwin_1872_01
all pro
darwin_1859_01 darwin_1860_01 darwin_1872_01
ce
darwin_1859_01 darwin_1860_01
n
darwin_1872_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
ded from a common parent, as any naturalist could in coming to a similar conclusion in regard to the many species of finches, or other
darwin_1859_01 darwin_1860_01
large
darwin_1859_01 darwin_1860_01 darwin_1872_01
groups of birds, in nature. One circumstance has struck me much; namely, that
darwin_1872_01
nearly
darwin_1859_01 darwin_1860_01 darwin_1872_01
all the breeders of the various domestic animals and the cultivators of plants, with whom I have
darwin_1859_01 darwin_1860_01
ever
darwin_1859_01 darwin_1860_01 darwin_1872_01
conversed, or whose treatises I have read, are firmly convinced that the several breeds to which each has attended, are descended from so many aboriginally distinct species. Ask, as I have asked, a celebrated raiser of Hereford cattle, whether his cattle might not have descended from
darwin_1859_01 darwin_1860_01
l
darwin_1872_01
L
darwin_1859_01 darwin_1860_01 darwin_1872_01
ong-horns,
darwin_1872_01
or both from a common parent-stock,
darwin_1859_01 darwin_1860_01 darwin_1872_01
and he will laugh you to scorn. I have never met a pigeon, or poultry, or duck, or rabbit fancier, who was not fully convinced that each main breed was descended from a distinct species. Van Mons, in his treatise on pears and apples, shows how utterly he disbelieves that the several sorts, for instance a Ribston-pippin or Codlin-apple, could ever have proceeded from the seeds of the same tree. Innumerable other examples could be given. The explanation, I think, is simple: from long-continued study they are strongly impressed with the differences between the several races; and though they well know that each race varies slightly, for they win their prizes by selecting such slight differences, yet they ignore all general arguments, and refuse to sum up in their minds slight differences accumulated during many successive generations. May not those naturalists who, knowing far less of the laws of inheritance than does the breeder, and knowing no more than he does of the intermediate links in the long lines of descent, yet admit that many of our domestic races
darwin_1859_01 darwin_1860_01
h
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
v
darwin_1872_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
e descended from the same parents—may they not learn a lesson of caution, when they deride the idea of species in a state of nature being lineal descendants of other species?
darwin_1872_01
Principles of
darwin_1859_01 darwin_1860_01 darwin_1872_01
Selection
darwin_1872_01
anciently followed, and their Effects
darwin_1859_01 darwin_1860_01 darwin_1872_01
.
darwin_1859_01 darwin_1860_01
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
Let us now briefly consider the steps by which domestic races have been produced, either from one or from several allied species. Some
darwin_1859_01 darwin_1860_01
little
darwin_1859_01 darwin_1860_01 darwin_1872_01
effect may
darwin_1859_01 darwin_1860_01
, perhaps,
darwin_1859_01 darwin_1860_01 darwin_1872_01
be attributed to the direct
darwin_1872_01
and definite
darwin_1859_01 darwin_1860_01 darwin_1872_01
action of the external conditions of life, and some
darwin_1859_01 darwin_1860_01
little
darwin_1859_01 darwin_1860_01 darwin_1872_01
to habit; but he would be a bold man who would account by such agencies for the differences
darwin_1859_01 darwin_1860_01
of
darwin_1872_01
between
darwin_1859_01 darwin_1860_01 darwin_1872_01
a dray and race horse, a greyhound and bloodhound, a carrier and tumbler pigeon. One of the most remarkable features in our domesticated races is that we see in them adaptation, not indeed to the animal's or plant's own good, but to man's use or fancy. Some variations useful to him have probably arisen suddenly, or by one step; many botanists, for instance, believe that the fuller's tea
darwin_1859_01 darwin_1860_01
zl
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
l
darwin_1859_01 darwin_1860_01 darwin_1872_01
, with its hooks, which cannot be rivalled by any mechanical contrivance, is only a variety of the wild Dipsacus; and this amount of change may have suddenly arisen in a seedling. So it has probably been with the turnspit dog; and this is known to have been the case with the ancon sheep. But when we compare the dray-horse and race-horse, the dromedary and camel, the various breeds of sheep fitted either for cultivated land or mountain pasture, with the wool of one breed good for one purpose, and that of another breed for another purpose; when we compare the many breeds of dogs, each good for man in
darwin_1859_01 darwin_1860_01
very
darwin_1859_01 darwin_1860_01 darwin_1872_01
different ways; when we compare the game-cock, so pertinacious in battle, with other breeds so little quarrel
darwin_1860_01
-
darwin_1859_01 darwin_1860_01 darwin_1872_01
some, with "everlasting layers" which never desire to sit, and with the bantam so small and elegant; when we compare the host of agricultural, culinary, orchard, and flower-garden races of plants, most useful to man at different seasons and for different purposes, or so beautiful in his eyes, we must, I think, look further than to mere variability. We cannot suppose that all the breeds were suddenly produced as perfect and as useful as we now see them; indeed, in
darwin_1859_01 darwin_1860_01
sever
darwin_1872_01
m
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
l
darwin_1872_01
ny
darwin_1859_01 darwin_1860_01 darwin_1872_01
cases, we know that this has not been their history. The key is man's power of accumulative selection: nature gives successive variations; man adds them up in certain directions useful to him. In this sense he may be said to
darwin_1872_01
have
darwin_1859_01 darwin_1860_01 darwin_1872_01
ma
darwin_1859_01 darwin_1860_01
k
darwin_1872_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
e for himself useful breeds. The great power of this principle of selection is not hypothetical. It is certain that several of our eminent breeders have, even within a single lifetime, modified to a large extent
darwin_1859_01 darwin_1860_01
som
darwin_1872_01
th
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
ir
darwin_1859_01 darwin_1860_01 darwin_1872_01
breeds of cattle and sheep. In order fully to realise what they have done, it is almost necessary to read several of the many treatises devoted to this subject, and to inspect the animals. Breeders habitually speak of an animal's organisation as something
darwin_1859_01 darwin_1860_01
quite
darwin_1859_01 darwin_1860_01 darwin_1872_01
plastic, which they can model almost as they please. If I had space I could quote numerous passages to this effect from highly competent authorities. Youatt, who was probably better acquainted with the works of agricultur
darwin_1859_01
al
darwin_1859_01 darwin_1860_01 darwin_1872_01
ists than almost any other individual, and who was himself a very good judge of an
darwin_1859_01 darwin_1860_01
an
darwin_1859_01 darwin_1860_01 darwin_1872_01
imal
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
, speaks of the principle of selection as "that which enables the agriculturist, not only to modify the character of his flock, but to change it altogether. It is the magician's wand, by means of which he may summon into life whatever form and mould he pleases." Lord Somerville, speaking of what breeders have done for sheep, says:—"It would seem as if they had chalked out upon a wall a form perfect in itself, and then had given it existence."
darwin_1859_01 darwin_1860_01
That most skilful breeder, Sir John Sebright, used to say, with respect to pigeons, that "he would produce any given feather in three years, but it would take him six years to obtain head and beak."
darwin_1859_01 darwin_1860_01 darwin_1872_01
In Saxony the importance of the principle of selection in regard to merino sheep is so fully recognised, that men follow it as a trade
darwin_1859_01 darwin_1860_01
:
darwin_1872_01
;
darwin_1859_01 darwin_1860_01 darwin_1872_01
the sheep are placed on a table and are studied, like a picture by a connoisseur; this is done three times at intervals of months, and the sheep are each time marked and classed, so that the very best may ultimately be selected for breeding. What English breeders have actually effected is proved by the enormous prices given for animals with a good pedigree; and these have
darwin_1859_01 darwin_1860_01
now
darwin_1859_01 darwin_1860_01 darwin_1872_01
been exported to almost every quarter of the world. The improvement is by no means generally due to crossing different breeds; all the best breeders are strongly opposed to this practice, except sometimes amongst closely allied sub-breeds. And when a cross has been made, the closest selection is far more indispensable even than in ordinary cases. If selection consisted merely in separating some very distinct variety, and breeding from it, the principle would be so obvious as hardly to be worth notice; but its importance consists in the great effect produced by the accumulation in one direction, during successive generations, of differences absolutely inappreciable by an uneducated eye—differences which I for one have vainly attempted to appreciate. Not one man in a thousand has accuracy of eye and judgment sufficient to become an eminent breeder. If gifted with these qualities, and he studies his subject for years, and devotes his lifetime to it with indomitable perseverance, he will succeed, and may make great improvements; if he wants any of these qualities, he will assuredly fail. Few would readily believe in the natural capacity and years of practice requisite to become even a skilful pigeon-fancier. The same principles are followed by horticulturists; but the variations are here often more abrupt. No one supposes that our choicest productions have been produced by a single variation from the aboriginal stock. We have proofs that this
darwin_1859_01 darwin_1860_01
i
darwin_1872_01
ha
darwin_1859_01 darwin_1860_01 darwin_1872_01
s not
darwin_1872_01
been
darwin_1859_01 darwin_1860_01 darwin_1872_01
so in s
darwin_1859_01 darwin_1860_01
om
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
veral
darwin_1859_01 darwin_1860_01 darwin_1872_01
cases
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
in which exact records have been kept; thus, to give a very trifling instance, the steadily-increasing size of the common gooseberry may be quoted. We see an astonishing improvement in many florists' flowers, when the flowers of the present day are compared with drawings made only twenty or thirty years ago. When a race of plants is once pretty well established, the seed-raisers do not pick out the best plants, but merely go over their seed-beds, and pull up the "rogues," as they call the plants that deviate from the proper standard. With animals this kind of selection is, in fact,
darwin_1859_01 darwin_1860_01
a
darwin_1859_01 darwin_1860_01 darwin_1872_01
l
darwin_1872_01
ikewi
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1859_01 darwin_1860_01
o
darwin_1872_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
followed; for hardly any one is so careless as to
darwin_1859_01 darwin_1860_01
all
darwin_1872_01
breed fr
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
w
darwin_1872_01
m
darwin_1859_01 darwin_1860_01 darwin_1872_01
his worst animals
darwin_1859_01 darwin_1860_01
to breed
darwin_1859_01 darwin_1860_01 darwin_1872_01
. In regard to plants, there is another means of observing the accumulated effects of selection—namely, by comparing the diversity of flowers in the different varieties of the same species in the flower-garden; the diversity of leaves, pods, or tubers, or whatever part is valued, in the kitchen-garden, in comparison with the flowers of the same varieties; and the diversity of fruit of the same species in the orchard, in comparison with the leaves and flowers of the same set of varieties. See how different the leaves of the cabbage are, and how extremely alike the flowers; how unlike the flowers of the heartsease are, and how alike the leaves; how much the fruit of the different kinds of gooseberries differ in size, colour, shape, and hairiness, and yet the flowers present very slight differences. It is not that the varieties which differ largely in some one point do not differ at all in other points; this is hardly ever,
darwin_1872_01
—I
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1872_01
speak after careful observation,—
darwin_1859_01 darwin_1860_01 darwin_1872_01
perhaps never, the case. The law
darwin_1859_01 darwin_1860_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
of correlat
darwin_1872_01
ed variat
darwin_1859_01 darwin_1860_01 darwin_1872_01
ion
darwin_1859_01 darwin_1860_01
of growth
darwin_1859_01 darwin_1860_01 darwin_1872_01
, the importance of which should never be overlooked, will ensure some differences; but, as a general rule,
darwin_1859_01 darwin_1860_01
I
darwin_1872_01
it
darwin_1859_01 darwin_1860_01 darwin_1872_01
cannot
darwin_1872_01
be
darwin_1859_01 darwin_1860_01 darwin_1872_01
doubt
darwin_1872_01
ed
darwin_1859_01 darwin_1860_01 darwin_1872_01
that the continued selection of slight variations, either in the leaves, the flowers, or the fruit, will produce races differing from each other chiefly in these characters. It may be objected that the principle of selection has been reduced to methodical practice for scarcely more than three-quarters of a century; it has certainly been more attended to of late years, and many treatises have been published on the subject; and the result
darwin_1859_01
, I may add,
darwin_1859_01 darwin_1860_01 darwin_1872_01
has been, in a corresponding degree, rapid and important. But it is very far from true that the principle is a modern discovery. I could give several references to
darwin_1872_01
works of high antiquity, in which
darwin_1859_01 darwin_1860_01 darwin_1872_01
the full
darwin_1859_01 darwin_1860_01
acknowledgment of the
darwin_1859_01 darwin_1860_01 darwin_1872_01
importance of the principle i
darwin_1859_01 darwin_1860_01
n work
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1859_01 darwin_1860_01
of high
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1872_01
ck
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1859_01 darwin_1860_01
tiquity
darwin_1872_01
owledged
darwin_1859_01 darwin_1860_01 darwin_1872_01
. In rude and barbarous periods of English history choice animals were often imported, and laws were passed to prevent their exportation: the destruction of horses under a certain size was ordered, and this may be compared to the "roguing" of plants by nurserymen. The principle of selection I find distinctly given in an ancient Chinese encyclopædia. Explicit rules are laid down by some of the Roman classical writers. From passages in Genesis, it is clear that the colour of domestic animals was at that early period attended to. Savages now sometimes cross their dogs with wild canine animals, to improve the breed, and they formerly did so, as is attested by passages in Pliny. The savages in South Africa match their draught cattle by colour, as do some of the Esquimaux their teams of dogs. Livingstone s
darwin_1859_01 darwin_1860_01
how
darwin_1872_01
tate
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1872_01
t
darwin_1859_01 darwin_1860_01 darwin_1872_01
h
darwin_1859_01 darwin_1860_01
ow much
darwin_1872_01
at
darwin_1859_01 darwin_1860_01 darwin_1872_01
good domestic breeds are
darwin_1872_01
highly
darwin_1859_01 darwin_1860_01 darwin_1872_01
valued by the negroes
darwin_1859_01 darwin_1860_01
of
darwin_1872_01
in
darwin_1859_01 darwin_1860_01 darwin_1872_01
the interior of Africa who have not associated with Europeans. Some of these facts do not show actual selection, but they show that the breeding of domestic animals was carefully attended to in ancient times, and is now attended to by the lowest savages. It would, indeed, have been a strange fact, had attention not been paid to breeding, for the inheritance of good and bad qualities is so obvious.
darwin_1872_01
Unconscious Selection.
darwin_1859_01 darwin_1860_01 darwin_1872_01
At the present time, eminent breeders try by methodical selection, with a distinct object in view, to make a new strain or sub-breed, superior to anything
darwin_1872_01
of th
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
xist
darwin_1872_01
k
darwin_1859_01 darwin_1860_01 darwin_1872_01
in
darwin_1859_01 darwin_1860_01
g
darwin_1872_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
in the country. But, for our purpose, a
darwin_1859_01 darwin_1860_01
kind
darwin_1872_01
form
darwin_1859_01 darwin_1860_01 darwin_1872_01
of Selection, which may be called Unconscious, and which results from every one trying to possess and breed from the best individual animals, is more important. Thus, a man who intends keeping pointers naturally tries to get as good dogs as he can, and afterwards breeds from his own best dogs, but he has no wish or expectation of permanently altering the breed. Nevertheless
darwin_1859_01 darwin_1860_01
I
darwin_1872_01
we
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
c
darwin_1872_01
m
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
nnot
darwin_1872_01
y
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
doubt
darwin_1872_01
infer
darwin_1859_01 darwin_1860_01 darwin_1872_01
that this process, continued during centuries, would improve and modify any breed, in the same way as Bakewell, Collins, &c., by this very same process, only carried on more methodically, did greatly modify, even during their
darwin_1859_01 darwin_1860_01
own
darwin_1859_01 darwin_1860_01 darwin_1872_01
lifetimes, the forms and qualities of their cattle. Slow and insensible changes of this kind c
darwin_1859_01 darwin_1860_01
ould
darwin_1872_01
an
darwin_1859_01 darwin_1860_01 darwin_1872_01
never be recognised unless actual measurements or careful drawings of the breeds in question ha
darwin_1859_01 darwin_1860_01
d
darwin_1872_01
ve
darwin_1859_01 darwin_1860_01 darwin_1872_01
been made long ago, which m
darwin_1859_01 darwin_1860_01
ight
darwin_1872_01
ay
darwin_1859_01 darwin_1860_01 darwin_1872_01
serve for comparison. In some cases, however, unchanged
darwin_1860_01 darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
or but little changed individuals of the same breed
darwin_1859_01 darwin_1860_01
may b
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
found
darwin_1872_01
xist
darwin_1859_01 darwin_1860_01 darwin_1872_01
in less civilised districts, where the breed has been less improved. There is reason to believe that King Charles's spaniel has been unconsciously modified to a large extent since the time of that monarch. Some highly competent authorities are convinced that the setter is directly derived from the spaniel, and has probably been slowly altered from it. It is known that the English pointer has been greatly changed within the last century, and in this case the change has, it is believed, been chiefly effected by crosses with the fox
darwin_1859_01 darwin_1860_01
-
darwin_1859_01 darwin_1860_01 darwin_1872_01
hound; but what concerns us is, that the change has been effected unconsciously and gradually, and yet so effectually, that, though the old Spanish pointer certainly came from Spain, Mr. Borrow has not seen, as I am informed by him, any native dog in Spain like our pointer. By a similar process of selection, and by careful training,
darwin_1859_01 darwin_1860_01
the whole body of
darwin_1859_01 darwin_1860_01 darwin_1872_01
English racehorses have come to surpass in fleetness and size the parent Arab
darwin_1859_01 darwin_1860_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1859_01 darwin_1860_01
tock
darwin_1859_01 darwin_1860_01 darwin_1872_01
, so that the latter, by the regulations for the Goodwood Races, are favoured in the weights
darwin_1872_01
which
darwin_1859_01 darwin_1860_01 darwin_1872_01
they carry. Lord Spencer and others have shown how the cattle of England have increased in weight and in early maturity, compared with the stock formerly kept in this country. By comparing the accounts given in
darwin_1872_01
various
darwin_1859_01 darwin_1860_01 darwin_1872_01
old
darwin_1859_01 darwin_1860_01
pigeon
darwin_1859_01 darwin_1860_01 darwin_1872_01
treatises of
darwin_1872_01
the former and present state of
darwin_1859_01 darwin_1860_01 darwin_1872_01
carrier
darwin_1859_01 darwin_1860_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
and tumbler
darwin_1859_01 darwin_1860_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
w
darwin_1872_01
p
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
th these breeds as now existin
darwin_1859_01 darwin_1860_01 darwin_1872_01
g
darwin_1872_01
eons
darwin_1859_01 darwin_1860_01 darwin_1872_01
in Britain, India, and Persia, we can
darwin_1859_01 darwin_1860_01
, I think, clearly
darwin_1859_01 darwin_1860_01 darwin_1872_01
trace the stages through which they have insensibly passed, and come to differ so greatly from the rock-pigeon. Youatt gives an excellent illustration of the effects of a course of selection, which may be considered as unconscious
darwin_1859_01 darwin_1860_01
ly followed
darwin_1859_01 darwin_1860_01 darwin_1872_01
, in so far that the breeders could never have expected
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
or even
darwin_1859_01 darwin_1860_01
have
darwin_1859_01 darwin_1860_01 darwin_1872_01
wished
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
to
darwin_1859_01 darwin_1860_01
have
darwin_1859_01 darwin_1860_01 darwin_1872_01
produce
darwin_1859_01 darwin_1860_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
the result which ensued—namely, the production of two distinct strains. The two flocks of Leicester sheep kept by Mr. Buckley and Mr. Burgess, as Mr. Youatt remarks, "have been purely bred from the original stock of Mr. Bakewell for upwards of fifty years. There is not a suspicion existing in the mind of any one at all acquainted with the subject
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
that the owner of either of them has deviated in any one instance from the pure blood of Mr. Bakewell
darwin_1859_01 darwin_1872_01
'
darwin_1859_01 darwin_1860_01 darwin_1872_01
s flock, and yet the difference between the sheep possessed by these two gentlemen is so great that they have the appearance of being quite different varieties." If there exist savages so barbarous as never to think of the inherited character of the offspring of their domestic animals, yet any one animal particularly useful to them, for any special purpose, would be carefully preserved during famines and other accidents, to which savages are so liable, and such choice animals would thus generally leave more offspring than the inferior ones; so that in this case there would be a kind of unconscious selection going on. We see the value set on animals even by the barbarians of Tierra del Fuego, by their killing and devouring their old women, in times of dearth, as of less value than their dogs. In plants the same gradual process of improvement, through the occasional preservation of the best individuals, whether or not sufficiently distinct to be ranked at their first appearance as distinct varieties, and whether or not two or more species or races have become blended together by crossing, may plainly be recognised in the increased size and beauty which we now see in the varieties of the heartsease, rose, pelargonium, dahlia, and other plants, when compared with the older varieties or with their parent-stocks. No one would ever expect to get a first-rate heartsease or dahlia from the seed of a wild plant. No one would expect to raise a first-rate melting pear from the seed of the wild pear, though he might succeed from a poor seedling growing wild, if it had come from a garden-stock. The pear, though cultivated in classical times, appears, from Pliny's description, to have been a fruit of very inferior quality. I have seen great surprise expressed in horticultural works at the wonderful skill of gardeners, in having produced such splendid results from such poor materials; but the art
darwin_1859_01 darwin_1860_01
, I cannot doubt,
darwin_1859_01 darwin_1860_01 darwin_1872_01
has been simple, and, as far as the final result is concerned, has been followed almost unconsciously. It has consisted in always cultivating the best known variety, sowing its seeds, and, when a slightly better variety
darwin_1859_01 darwin_1860_01
has
darwin_1859_01 darwin_1860_01 darwin_1872_01
chanced to appear, selecting it, and so onwards. But the gardeners of the classical period, who cultivated the best pear
darwin_1872_01
s which
darwin_1859_01 darwin_1860_01 darwin_1872_01
they could procure, never thought what splendid fruit we should eat; though we owe our excellent fruit, in some small degree, to their having naturally chosen and preserved the best varieties they could anywhere find. A large amount of change
darwin_1859_01 darwin_1860_01
in our cultivated plants
darwin_1859_01 darwin_1860_01 darwin_1872_01
, thus slowly and unconsciously accumulated, explains, as I believe, the well-known fact, that in a
darwin_1859_01 darwin_1860_01
vast
darwin_1859_01 darwin_1860_01 darwin_1872_01
number of cases we cannot recognise, and therefore do not know, the wild parent-stocks of the plants which have been longest cultivated in our flower and kitchen gardens. If it has taken centuries or thousands of years to improve or modify most of our plants up to their present standard of usefulness to man, we can understand how it is that neither Australia, the Cape of Good Hope, nor any other region inhabited by quite uncivilised man, has afforded us a single plant worth culture. It is not that these countries, so rich in species, do not by a strange chance possess the aboriginal stocks of any useful plants, but that the native plants have not been improved by continued selection up to a standard of perfection comparable with that
darwin_1859_01 darwin_1860_01
g
darwin_1872_01
acqu
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
v
darwin_1872_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
n
darwin_1872_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
to
darwin_1872_01
by
darwin_1859_01 darwin_1860_01 darwin_1872_01
the plants in countries anciently civilised. In regard to the domestic animals kept by uncivilised man, it should not be overlooked that they almost always have to struggle for their own food, at least during certain seasons. And in two countries very differently circumstanced, individuals of the same species, having slightly different constitutions or structure, would often succeed better in the one country than in the other
darwin_1860_01 darwin_1872_01
;
darwin_1859_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
and thus by a process of "natural selection," as will hereafter be more fully explained, two sub-breeds might be formed. This, perhaps, partly explains wh
darwin_1859_01 darwin_1860_01
at has been remarked b
darwin_1859_01 darwin_1860_01 darwin_1872_01
y
darwin_1859_01 darwin_1860_01
some authors, namely, that
darwin_1859_01 darwin_1860_01 darwin_1872_01
the varieties kept by savages
darwin_1872_01
, as has been remarked by some authors,
darwin_1859_01 darwin_1860_01 darwin_1872_01
have more of the character of
darwin_1872_01
true
darwin_1859_01 darwin_1860_01 darwin_1872_01
species than the varieties kept in civilised countries. On the view here given of the
darwin_1859_01 darwin_1860_01
all-
darwin_1859_01 darwin_1860_01 darwin_1872_01
important part which selection by man has played, it becomes at once obvious, how it is that our domestic races show adaptation in their structure or in their habits to man
darwin_1859_01 darwin_1872_01
'
darwin_1859_01 darwin_1860_01 darwin_1872_01
s wants or fancies. We can, I think, further understand the frequently abnormal character of our domestic races, and likewise their differences being so great in external characters
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
and relatively so slight in internal parts or organs. Man can hardly select, or only with much difficulty, any deviation of structure excepting such as is externally visible; and indeed he rarely cares for what is internal. He can never act by selection, excepting on variations
darwin_1859_01 darwin_1872_01
which are first given to him in some slight degree by nature. No man would ever try to make a
darwin_1859_01 darwin_1860_01 darwin_1872_01
fantail
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
till he saw a pigeon with a tail developed in some slight degree in an unusual manner, or a pouter till he saw a pigeon with a crop of somewhat unusual size; and the more abnormal or unusual any character was when it first appeared, the more likely it would be to catch his attention. But to use such an expression as trying to make a fantail, is, I have no doubt, in most cases, utterly incorrect. The man who first selected a pigeon with a slightly larger tail, never dreamed what the descendants of that pigeon would become through long-continued, partly unconscious and partly methodical
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
selection. Perhaps the parent
darwin_1859_01 darwin_1860_01
darwin_1872_01
-
darwin_1859_01 darwin_1860_01 darwin_1872_01
bird of all fantails had only fourteen tail-feathers somewhat expanded, like the present Java fantail, or like individuals of other and distinct breeds, in which as many as seventeen tail-feathers have been counted. Perhaps the first pouter-pigeon did not inflate its crop much more than the turbit now does the upper part of its œsophagus,—a habit which is disregarded by all fanciers, as it is not one of the points of the breed. Nor let it be thought that some great deviation of structure would be necessary to catch the fancier
darwin_1859_01 darwin_1872_01
'
darwin_1859_01 darwin_1860_01 darwin_1872_01
s eye: he perceives extremely small differences, and it is in human nature to value any novelty, however slight, in one
darwin_1859_01 darwin_1872_01
'
darwin_1859_01 darwin_1860_01 darwin_1872_01
s own possession. Nor must the value which would formerly
darwin_1872_01
have
darwin_1859_01 darwin_1860_01 darwin_1872_01
be
darwin_1872_01
en
darwin_1859_01 darwin_1860_01 darwin_1872_01
set on any slight differences in the individuals of the same species, be judged of by the value which
darwin_1859_01 darwin_1860_01
would
darwin_1872_01
is
darwin_1859_01 darwin_1860_01 darwin_1872_01
now
darwin_1859_01 darwin_1860_01
be
darwin_1859_01 darwin_1860_01 darwin_1872_01
set on them, after several breeds have
darwin_1859_01 darwin_1860_01
once
darwin_1859_01 darwin_1860_01 darwin_1872_01
fairly been established.
darwin_1859_01 darwin_1860_01
M
darwin_1872_01
It is known that with pigeons m
darwin_1859_01 darwin_1860_01 darwin_1872_01
any slight
darwin_1859_01 darwin_1860_01
differences might,
darwin_1872_01
v
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
nd
darwin_1872_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
ndeed d
darwin_1872_01
ati
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1872_01
ns
darwin_1859_01 darwin_1860_01 darwin_1872_01
now
darwin_1872_01
occasionally appear
darwin_1859_01 darwin_1860_01 darwin_1872_01
,
darwin_1859_01 darwin_1860_01
ari
darwin_1872_01
but the
darwin_1859_01 darwin_1860_01 darwin_1872_01
se
darwin_1859_01 darwin_1860_01
amongst pigeons, which
darwin_1859_01 darwin_1860_01 darwin_1872_01
are rejected as faults or deviations from the standard of perfection
darwin_1859_01 darwin_1860_01
of
darwin_1872_01
in
darwin_1859_01 darwin_1860_01 darwin_1872_01
each breed. The common goose has not given rise to any marked varieties; hence the T
darwin_1859_01 darwin_1860_01
h
darwin_1859_01 darwin_1860_01 darwin_1872_01
oulouse and the common breed, which differ only in colour, that most fleeting of characters, have lately been exhibited as distinct at our poultry-shows.
darwin_1859_01 darwin_1860_01
I think t
darwin_1872_01
T
darwin_1859_01 darwin_1860_01 darwin_1872_01
hese views
darwin_1859_01 darwin_1860_01
fu
darwin_1872_01
appea
darwin_1859_01 darwin_1860_01 darwin_1872_01
r
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
her
darwin_1872_01
o
darwin_1859_01 darwin_1860_01 darwin_1872_01
explain what has sometimes been noticed—namely
darwin_1860_01 darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
that we know
darwin_1872_01
hardly a
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1859_01 darwin_1860_01
o
darwin_1872_01
y
darwin_1859_01 darwin_1860_01 darwin_1872_01
thing about the origin or history of any of our domestic breeds. But, in fact, a breed, like a dialect of a language, can hardly be said to have
darwin_1859_01 darwin_1860_01
had
darwin_1859_01 darwin_1860_01 darwin_1872_01
a d
darwin_1859_01 darwin_1860_01
ef
darwin_1872_01
ist
darwin_1859_01 darwin_1860_01 darwin_1872_01
in
darwin_1859_01 darwin_1860_01
i
darwin_1872_01
c
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
origin. A man preserves and breeds from an individual with some slight deviation of structure, or takes more care than usual in matching his best animals
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
and thus improves them, and the improved
darwin_1859_01 darwin_1860_01
i
darwin_1872_01
a
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1859_01 darwin_1860_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
vidu
darwin_1872_01
m
darwin_1859_01 darwin_1860_01 darwin_1872_01
als slowly spread in the immediate neighbourhood. But
darwin_1859_01 darwin_1860_01
as yet
darwin_1859_01 darwin_1860_01 darwin_1872_01
they will
darwin_1872_01
as yet
darwin_1859_01 darwin_1860_01 darwin_1872_01
hardly have a distinct name, and from being only slightly valued, their history will
darwin_1872_01
have
darwin_1859_01 darwin_1860_01 darwin_1872_01
be
darwin_1872_01
en
darwin_1859_01 darwin_1860_01 darwin_1872_01
disregarded. When further improved by the same slow and gradual process, they will spread more widely, and will
darwin_1859_01 darwin_1860_01
g
darwin_1872_01
b
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
t
darwin_1859_01 darwin_1860_01 darwin_1872_01
recognised as something distinct and valuable, and will then probably first receive a provincial name. In semi-civilised countries, with little free communication, the spreading
darwin_1859_01 darwin_1860_01
and knowledge
darwin_1859_01 darwin_1860_01 darwin_1872_01
of a
darwin_1859_01 darwin_1860_01
ny
darwin_1859_01 darwin_1860_01 darwin_1872_01
new sub-breed w
darwin_1859_01 darwin_1860_01
i
darwin_1872_01
ou
darwin_1859_01 darwin_1860_01 darwin_1872_01
l
darwin_1859_01 darwin_1860_01
l
darwin_1872_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
be a slow process. As soon as the points of value
darwin_1859_01 darwin_1860_01
of the new sub-breed
darwin_1859_01 darwin_1860_01 darwin_1872_01
are once
darwin_1859_01 darwin_1860_01
fully
darwin_1859_01 darwin_1860_01 darwin_1872_01
acknowledged, the principle, as I have called it, of unconscious selection will always tend,—
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
perhaps more at one period than at another, as the breed rises or falls in fashion,—perhaps more in one district than in another, according to the state of civili
darwin_1859_01 darwin_1872_01
s
darwin_1860_01
z
darwin_1859_01 darwin_1860_01 darwin_1872_01
ation of the inhabitants,—
darwin_1872_01
darwin_1859_01 darwin_1860_01 darwin_1872_01
slowly to add to the characteristic features of the breed, whatever they may be. But the chance will be infinitely small of any record having been preserved of such slow, varying, and insensible changes.
darwin_1872_01
Circumstances favourable to Man's Power of Selection.
darwin_1859_01 darwin_1860_01 darwin_1872_01
I
darwin_1859_01 darwin_1860_01
must
darwin_1872_01
will
darwin_1859_01 darwin_1860_01 darwin_1872_01
now say a few words on the circumstances, favourable, or the reverse, to man
darwin_1859_01 darwin_1872_01
'
darwin_1859_01 darwin_1860_01 darwin_1872_01
s power of selection. A high degree of variability is obviously favourable, as freely giving the materials for selection to work on; not that mere individual differences are not amply sufficient, with extreme care, to allow of the accumulation of a large amount of modification in almost any desired direction. But as variations manifestly useful or pleasing to man appear only occasionally, the chance of their appearance will be much increased by a large number of individuals being kept
darwin_1859_01 darwin_1860_01
;
darwin_1872_01
.
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
and h
darwin_1872_01
H
darwin_1859_01 darwin_1860_01 darwin_1872_01
ence
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
this co
darwin_1872_01
nu
darwin_1859_01 darwin_1860_01 darwin_1872_01
m
darwin_1859_01 darwin_1860_01
es to
darwin_1859_01 darwin_1860_01 darwin_1872_01
be
darwin_1872_01
r is
darwin_1859_01 darwin_1860_01 darwin_1872_01
of the highest importance
darwin_1859_01 darwin_1860_01
t
darwin_1872_01
f
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1872_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
success. On this principle Marshall
darwin_1859_01 darwin_1860_01
has
darwin_1872_01
formerly
darwin_1859_01 darwin_1860_01 darwin_1872_01
remarked, with respect to the sheep of parts of Yorkshire,
darwin_1859_01 darwin_1860_01
that
darwin_1859_01 darwin_1860_01 darwin_1872_01
"as they generally belong to poor people, and are mostly in small lots, they never can be improved." On the other hand, nurserymen, from
darwin_1859_01 darwin_1860_01
rais
darwin_1872_01
keep
darwin_1859_01 darwin_1860_01 darwin_1872_01
ing large stocks of the same plant
darwin_1859_01 darwin_1860_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
, are generally far more successful than amateurs in
darwin_1859_01 darwin_1860_01
gett
darwin_1872_01
rais
darwin_1859_01 darwin_1860_01 darwin_1872_01
ing new and valuable varieties.
darwin_1859_01 darwin_1860_01
The keeping of a
darwin_1872_01
A
darwin_1859_01 darwin_1860_01 darwin_1872_01
large number of individuals of a
darwin_1859_01 darwin_1860_01
species i
darwin_1859_01 darwin_1860_01 darwin_1872_01
n an
darwin_1859_01 darwin_1860_01
y country requ
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
res th
darwin_1872_01
m
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
t the species shou
darwin_1859_01 darwin_1860_01 darwin_1872_01
l
darwin_1859_01 darwin_1860_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
be
darwin_1872_01
or
darwin_1859_01 darwin_1860_01 darwin_1872_01
pla
darwin_1872_01
nt
darwin_1859_01 darwin_1860_01 darwin_1872_01
c
darwin_1872_01
an be rear
darwin_1859_01 darwin_1860_01 darwin_1872_01
ed
darwin_1859_01 darwin_1860_01
u
darwin_1872_01
o
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1859_01 darwin_1860_01
d
darwin_1872_01
ly wh
darwin_1859_01 darwin_1860_01 darwin_1872_01
er
darwin_1872_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
favourabl
darwin_1872_01
th
darwin_1859_01 darwin_1860_01 darwin_1872_01
e conditions
darwin_1859_01 darwin_1860_01
o
darwin_1859_01 darwin_1860_01 darwin_1872_01
f
darwin_1859_01 darwin_1860_01
life, s
darwin_1859_01 darwin_1860_01 darwin_1872_01
o
darwin_1859_01 darwin_1860_01
as to b
darwin_1859_01 darwin_1860_01 darwin_1872_01
r
darwin_1859_01 darwin_1860_01
eed freely
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1872_01
ts propagatio
darwin_1859_01 darwin_1860_01 darwin_1872_01
n
darwin_1859_01 darwin_1860_01
th
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1859_01 darwin_1860_01
t
darwin_1872_01
re
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
c
darwin_1872_01
fav
darwin_1859_01 darwin_1860_01 darwin_1872_01
ou
darwin_1859_01 darwin_1860_01
nt
darwin_1859_01 darwin_1860_01 darwin_1872_01
r
darwin_1859_01 darwin_1860_01
y
darwin_1872_01
able
darwin_1859_01 darwin_1860_01 darwin_1872_01
. When the individuals
darwin_1859_01 darwin_1860_01
of any species
darwin_1859_01 darwin_1860_01 darwin_1872_01
are scanty, all
darwin_1859_01 darwin_1860_01
th
darwin_1872_01
will b
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
individu
darwin_1859_01 darwin_1860_01 darwin_1872_01
al
darwin_1859_01 darwin_1860_01
s
darwin_1872_01
lowed to breed
darwin_1859_01 darwin_1860_01 darwin_1872_01
, whatever their quality may be,
darwin_1859_01 darwin_1860_01
will generally be allowed to breed,
darwin_1859_01 darwin_1860_01 darwin_1872_01
and this will effectually prevent selection. But probably the most important
darwin_1859_01 darwin_1860_01
poi
darwin_1872_01
eleme
darwin_1859_01 darwin_1860_01 darwin_1872_01
nt
darwin_1859_01 darwin_1860_01
of all,
darwin_1859_01 darwin_1860_01 darwin_1872_01
is
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
that the animal or plant should be so highly
darwin_1859_01 darwin_1860_01
useful to man, or so much
darwin_1859_01 darwin_1860_01 darwin_1872_01
valued by
darwin_1859_01 darwin_1860_01
hi
darwin_1859_01 darwin_1860_01 darwin_1872_01
m
darwin_1872_01
an
darwin_1859_01 darwin_1860_01 darwin_1872_01
, that the closest attention
darwin_1872_01
i
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1859_01 darwin_1860_01
hould be
darwin_1859_01 darwin_1860_01 darwin_1872_01
paid to even the slightest deviation
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
in
darwin_1872_01
i
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
he
darwin_1872_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
qualities or structure
darwin_1859_01 darwin_1860_01
of each individual
darwin_1859_01 darwin_1860_01 darwin_1872_01
. Unless such attention be paid nothing can be effected. I have seen it gravely remarked, that it was most fortunate that the strawberry began to vary just when gardeners began to attend
darwin_1859_01 darwin_1860_01
closely
darwin_1859_01 darwin_1860_01 darwin_1872_01
to this plant. No doubt the strawberry had always varied since it was cultivated, but the slight varieties had been neglected. As soon, however, as gardeners picked out individual plants with slightly larger, earlier, or better fruit, and raised seedlings from them, and again picked out the best seedlings and bred from them, then
darwin_1859_01 darwin_1860_01
, there appeared
darwin_1859_01 darwin_1860_01 darwin_1872_01
(
darwin_1859_01 darwin_1860_01
a
darwin_1872_01
w
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
ded by
darwin_1872_01
th
darwin_1859_01 darwin_1860_01 darwin_1872_01
some
darwin_1872_01
aid by
darwin_1859_01 darwin_1860_01 darwin_1872_01
crossing
darwin_1859_01 darwin_1860_01
with
darwin_1859_01 darwin_1860_01 darwin_1872_01
distinct species) those many admirable varieties of the strawberry w
darwin_1872_01
ere raised w
darwin_1859_01 darwin_1860_01 darwin_1872_01
hich have
darwin_1859_01 darwin_1860_01
b
darwin_1872_01
app
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1859_01 darwin_1860_01
en
darwin_1872_01
a
darwin_1859_01 darwin_1860_01 darwin_1872_01
r
darwin_1859_01 darwin_1860_01
ais
darwin_1859_01 darwin_1860_01 darwin_1872_01
ed during the last
darwin_1859_01 darwin_1860_01
t
darwin_1859_01 darwin_1860_01 darwin_1872_01
h
darwin_1859_01 darwin_1860_01
irty or forty ye
darwin_1859_01 darwin_1860_01 darwin_1872_01
a
darwin_1872_01
lf-centu
darwin_1859_01 darwin_1860_01 darwin_1872_01
r
darwin_1859_01 darwin_1860_01
s
darwin_1872_01
y
darwin_1859_01 darwin_1860_01 darwin_1872_01
.
darwin_1859_01 darwin_1860_01
In
darwin_1872_01
Wi
darwin_1859_01 darwin_1860_01 darwin_1872_01
th
darwin_1859_01 darwin_1860_01
e case of
darwin_1859_01 darwin_1860_01 darwin_1872_01
animal
darwin_1859_01 darwin_1860_01
s with separate sexe
darwin_1859_01 darwin_1860_01 darwin_1872_01
s, facility in preventing crosses is an important element
darwin_1859_01 darwin_1860_01
of success
darwin_1859_01 darwin_1860_01 darwin_1872_01
in the formation of new races,—at least, in a country which is already stocked with other races. In this respect enclosure of the land plays a part. Wandering savages or the inhabitants of open plains rarely possess more than one breed of the same species. Pigeons can be mated for life, and this is a great convenience to the fancier, for thus many races may be
darwin_1872_01
improved and
darwin_1859_01 darwin_1860_01 darwin_1872_01
kept true, though mingled in the same aviary; and this circumstance must have largely favoured the
darwin_1859_01 darwin_1860_01
improvement and
darwin_1859_01 darwin_1860_01 darwin_1872_01
formation of new breeds. Pigeons, I may add, can be propagated in great numbers and at a very quick rate, and inferior birds may be freely rejected, as when killed they serve for food. On the other hand, cats, from their nocturnal rambling habits, cannot be
darwin_1872_01
easily
darwin_1859_01 darwin_1860_01 darwin_1872_01
matched, and, although so much valued by women and children, we
darwin_1859_01 darwin_1860_01
h
darwin_1872_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
ar
darwin_1859_01 darwin_1860_01
d
darwin_1872_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
ly
darwin_1859_01 darwin_1860_01
ever
darwin_1859_01 darwin_1860_01 darwin_1872_01
see a distinct breed
darwin_1872_01
long
darwin_1859_01 darwin_1860_01 darwin_1872_01
kept up; such breeds as we do sometimes see are almost always imported from some other country
darwin_1859_01 darwin_1860_01
, often from islands
darwin_1859_01 darwin_1860_01 darwin_1872_01
. Although I do not doubt that some domestic animals vary less than others, yet the rarity or absence of distinct breeds of the cat, the donkey, peacock, goose, &c., may be attributed in main part to selection not having been brought into play: in cats, from the difficulty in pairing them; in donkeys, from only a few being kept by poor people, and little attention paid to their breeding;
darwin_1872_01
for recently in certain parts of Spain and of the United States this animal has been surprisingly modified and improved by careful selection;
darwin_1859_01 darwin_1860_01 darwin_1872_01
in peacocks, from not being very easily reared and a large stock not kept; in geese, from being valuable only for two purposes, food and feathers, and more especially from no pleasure having been felt in the display of distinct breeds
darwin_1872_01
; but the goose, under the conditions to which it is exposed when domesticated, seems to have a singularly inflexible organisation, though it has varied to a slight extent, as I have elsewhere described. Some authors have maintained that the amount of variation in our domestic productions is soon reached, and can never afterwards be exceeded. It would be somewhat rash to assert that the limit has been attained in any one case; for almost all our animals and plants have been greatly improved in many ways within a recent period; and this implies variation. It would be equally rash to assert that characters now increased to their utmost limit, could not, after remaining fixed for many centuries, again vary under new conditions of life. No doubt, as Mr. Wallace has remarked with much truth, a limit will be at last reached. For instance, there must be a limit to the fleetness of any terrestrial animal, as this will be determined by the friction to be overcome, the weight of body to be carried, and the power of contraction in the muscular fibres. But what concerns us is that the domestic varieties of the same species differ from each other in almost every character, which man has attended to and selected, more than do the distinct species of the same genera. Isidore Geoffroy St. Hilaire has proved this in regard to size, and so it is with colour and probably with the length of hair. With respect to fleetness, which depends on many bodily characters, Eclipse was far fleeter, and a dray-horse is incomparably stronger than any two natural species belonging to the same genus. So with plants, the seeds of the different varieties of the bean or maize probably differ more in size, than do the seeds of the distinct species in any one genus in the same two families. The same remark holds good in regard to the fruit of the several varieties of the plum, and still more strongly with the melon, as well as in many other analogous cases
darwin_1859_01 darwin_1860_01 darwin_1872_01
. To sum up on the origin of our
darwin_1859_01 darwin_1860_01
D
darwin_1872_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
omestic
darwin_1859_01 darwin_1860_01
R
darwin_1872_01
r
darwin_1859_01 darwin_1860_01 darwin_1872_01
aces of animals and plants.
darwin_1859_01 darwin_1860_01
I believe t
darwin_1872_01
C
darwin_1859_01 darwin_1860_01 darwin_1872_01
ha
darwin_1859_01 darwin_1860_01
t th
darwin_1872_01
ng
darwin_1859_01 darwin_1860_01 darwin_1872_01
e
darwin_1872_01
d
darwin_1859_01 darwin_1860_01 darwin_1872_01
conditions of life
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1872_01
are o
darwin_1859_01 darwin_1860_01 darwin_1872_01
f
darwin_1859_01 darwin_1860_01
rom
darwin_1859_01 darwin_1860_01 darwin_1872_01
the
darwin_1872_01
h
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1872_01
ghest impo
darwin_1859_01 darwin_1860_01 darwin_1872_01
r
darwin_1872_01
tance in causing variability, both by
darwin_1859_01 darwin_1860_01 darwin_1872_01
acti
darwin_1872_01
ng directly
darwin_1859_01 darwin_1860_01 darwin_1872_01
on
darwin_1872_01
the organisati
darwin_1859_01 darwin_1860_01 darwin_1872_01
on
darwin_1872_01
, and indirectly by affecting
darwin_1859_01 darwin_1860_01 darwin_1872_01
the reproductive system
darwin_1859_01 darwin_1860_01
, are so far of the highest importance as causing variability
darwin_1859_01 darwin_1860_01 darwin_1872_01
. I
darwin_1872_01
t
darwin_1859_01 darwin_1860_01 darwin_1872_01
darwin_1859_01 darwin_1860_01
do
darwin_1872_01
is
darwin_1859_01 darwin_1860_01 darwin_1872_01
not
darwin_1872_01
pro
darwin_1859_01 darwin_1860_01 darwin_1872_01
b
darwin_1859_01 darwin_1860_01
e
darwin_1872_01
ab
darwin_1859_01 darwin_1860_01 darwin_1872_01
l
darwin_1859_01 darwin_1860_01
iev
darwin_1859_01 darwin_1860_01 darwin_1872_01
e that variability is an inherent and necessary contingen
darwin_1859_01 darwin_1860_01
cy
darwin_1872_01
t
darwin_1859_01 darwin_1860_01 darwin_1872_01
, under all circumstances
darwin_1859_01 darwin_1860_01
, with all organic beings, as some authors have thought
darwin_1859_01 darwin_1860_01 darwin_1872_01
. The
darwin_1859_01 darwin_1860_01
effects of variability are modified by various de
darwin_1859_01 darwin_1860_01 darwin_1872_01
gre
darwin_1872_01
ater or l
darwin_1859_01 darwin_1860_01 darwin_1872_01
es
darwin_1872_01
s force
darwin_1859_01 darwin_1860_01 darwin_1872_01
of inheritance and
darwin_1859_01 darwin_1860_01
of
darwin_1859_01 darwin_1860_01 darwin_1872_01
reversion
darwin_1872_01
determine whether variations shall endure
darwin_1859_01 darwin_1860_01 darwin_1872_01
. Variability is governed by many unknown laws,
darwin_1859_01 darwin_1860_01
more especially by that
darwin_1859_01 darwin_1860_01 darwin_1872_01
of
darwin_1872_01
which
darwin_1859_01 darwin_1860_01 darwin_1872_01
correlat
darwin_1859_01 darwin_1860_01
ion of
darwin_1872_01
ed
darwin_1859_01 darwin_1860_01 darwin_1872_01
growth
darwin_1872_01
is probably the most important
darwin_1859_01 darwin_1860_01 darwin_1872_01
. Something
darwin_1872_01
, but how much we do not know,
darwin_1859_01 darwin_1860_01 darwin_1872_01
may be attributed to the d
darwin_1872_01
ef
darwin_1859_01 darwin_1860_01 darwin_1872_01
i
darwin_1859_01 darwin_1860_01
rec
darwin_1872_01
ni
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1872_01
e
darwin_1859_01 darwin_1860_01 darwin_1872_01
action of the conditions of life. Some
darwin_1872_01
, perhaps a grea
darwin_1859_01 darwin_1860_01 darwin_1872_01
t
darwin_1859_01 darwin_1860_01
hing
darwin_1872_01
, effect
darwin_1859_01 darwin_1860_01 darwin_1872_01
m
darwin_1859_01 darwin_1860_01
ust
darwin_1872_01
ay
darwin_1859_01 darwin_1860_01 darwin_1872_01
be attributed to
darwin_1872_01
the[increased
darwin_1859_01 darwin_1860_01 darwin_1872_01
use
darwin_1859_01 darwin_1860_01
and
darwin_1872_01
or
darwin_1859_01 darwin_1860_01 darwin_1872_01
disuse
darwin_1872_01
of parts
darwin_1859_01 darwin_1860_01 darwin_1872_01
. The final result is thus rendered infinitely complex. In some cases
darwin_1859_01 darwin_1860_01
, I do not doubt that
darwin_1859_01 darwin_1860_01 darwin_1872_01
the intercrossing of
darwin_1859_01 darwin_1860_01
species,
darwin_1859_01 darwin_1860_01 darwin_1872_01
aboriginally distinct
darwin_1859_01 darwin_1860_01
,
darwin_1872_01
species appears to
darwin_1859_01 darwin_1860_01 darwin_1872_01
ha
darwin_1859_01 darwin_1860_01
s
darwin_1872_01
ve
darwin_1859_01 darwin_1860_01 darwin_1872_01
played an important part in the origin of our
darwin_1859_01 darwin_1860_01
domestic p
darwin_1872_01
b
darwin_1859_01 darwin_1860_01 darwin_1872_01
r
darwin_1859_01 darwin_1860_01
o
darwin_1872_01
ee
darwin_1859_01 darwin_1860_01 darwin_1872_01
d
darwin_1859_01 darwin_1860_01
uction
darwin_1859_01 darwin_1860_01 darwin_1872_01
s. When
darwin_1859_01 darwin_1860_01
in any country
darwin_1859_01 darwin_1860_01 darwin_1872_01
several
darwin_1859_01 darwin_1860_01
domestic
darwin_1859_01 darwin_1860_01 darwin_1872_01
breeds have once been
darwin_1859_01 darwin_1860_01
establish
darwin_1872_01
form
darwin_1859_01 darwin_1860_01 darwin_1872_01
ed
darwin_1872_01
in any country
darwin_1859_01 darwin_1860_01 darwin_1872_01
, their occasional intercrossing, with the aid of selection, has, no doubt, largely aided in the formation of new sub-breeds; but the importance of
darwin_1859_01 darwin_1860_01
the
darwin_1859_01 darwin_1860_01 darwin_1872_01
crossing
darwin_1859_01 darwin_1860_01
of varieties
darwin_1859_01 darwin_1860_01 darwin_1872_01
has
darwin_1859_01 darwin_1860_01
, I believe,
darwin_1859_01 darwin_1860_01 darwin_1872_01
been
darwin_1859_01 darwin_1860_01
greatly
darwin_1872_01
much
darwin_1859_01 darwin_1860_01 darwin_1872_01
exaggerated, both in regard to animals and to those plants which are propagated by seed.
darwin_1859_01 darwin_1860_01
In
darwin_1872_01
With
darwin_1859_01 darwin_1860_01 darwin_1872_01
plants which are temporarily propagated by cuttings, buds, &c., the importance of
darwin_1859_01 darwin_1860_01
the
darwin_1859_01 darwin_1860_01 darwin_1872_01
crossing
darwin_1859_01 darwin_1860_01
both of distinct species and of varieties
darwin_1859_01 darwin_1860_01 darwin_1872_01
is immense; for the cultivator
darwin_1872_01
may
darwin_1859_01 darwin_1860_01 darwin_1872_01
here
darwin_1859_01 darwin_1860_01
quite
darwin_1859_01 darwin_1860_01 darwin_1872_01
disregard
darwin_1859_01 darwin_1860_01
s
darwin_1859_01 darwin_1860_01 darwin_1872_01
the extreme variability both of hybrids and
darwin_1872_01
of
darwin_1859_01 darwin_1860_01 darwin_1872_01
mongrels, and the
darwin_1859_01 darwin_1860_01
frequent
darwin_1859_01 darwin_1860_01 darwin_1872_01
sterility of hybrids; but
darwin_1859_01 darwin_1860_01
the cases of
darwin_1859_01 darwin_1860_01 darwin_1872_01
plants not propagated by seed are of little importance to us, for their endurance is only temporary. Over all these causes of Change
darwin_1859_01 darwin_1860_01
I am convinced that
darwin_1872_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
the accumulative action of Selection, whether applied methodically and
darwin_1859_01 darwin_1860_01
more
darwin_1859_01 darwin_1860_01 darwin_1872_01
quickly, or unconsciously and
darwin_1859_01 darwin_1860_01
more
darwin_1859_01 darwin_1860_01 darwin_1872_01
slowly
darwin_1859_01 darwin_1860_01
,
darwin_1859_01 darwin_1860_01 darwin_1872_01
but more efficiently,
darwin_1859_01 darwin_1860_01
i
darwin_1872_01
seem
darwin_1859_01 darwin_1860_01 darwin_1872_01
s
darwin_1872_01
to have
darwin_1859_01 darwin_1860_01 darwin_1872_01
b
darwin_1859_01 darwin_1860_01
y far
darwin_1872_01
een
darwin_1859_01 darwin_1860_01 darwin_1872_01
the predominant Power.